Elizabeth B. Bryant.
Notes on North American Anyphaeninae in the Museum of Comparative Zoology.
Psyche 38(2-3):102-126, 1931.

This article at Hindawi Publishing: https://doi.org/10.1155/1931/13898
CEC's scan of this article: http://psyche.entclub.org/pdf/38/38-102.pdf, 1672K
This landing page: http://psyche.entclub.org/38/38-102.html

The following unprocessed text is extracted automatically from the PDF file, and is likely to be both incomplete and full of errors. Please consult the PDF file for the complete article.

Psyche
[ June-September
NOTES ON NORTH AMERICAN
ANYPHENINa
IN
THE MUSEUM OF COMPARATIVE ZOOLOGY
In a recent revision of the spiders of this sub-family in the Museum of Comparative Zoology from North and Cen- tral America and the West Indies, it was found that there has been much confusion in the use of genera. The genus Anyphsena was made by Sundevall in 1833 (Conspectus Arachnidum, p. 20) for Aranea accentuata Walckenaer. L. Koch (Die Arach. fam. Drassiden, 1866, p. 194) first called attention to the position of the ventral furrow, "as either in the middle of the length of the abdomen or behind the middle." Nicolet (Say, Hist. de Chili, 1849, 3, p. 450) made the genus Gayenna for the South American species where the posterior spiracular openings are nearer the spinnerets than to the epigastric plate. F. 0. P. Cambridge (Biologia Centrali-Americana, 1900, 2, p. 94), definitely restricts the genus to "Spiracular rima situated behind the middle of the ventral area of the abdomen towards the spinners," and states that the genus is not found in Central America. Mr. Banks has used the genus Gayenna when the ventral furrow is behind the middle of the entire abdomen. The genus Marcellina has been made for Clubiom pisca- toria Hentz, known only from two males from Alabama and an immature male from Newton, Massachusetts. The
large anterior median eyes, the peculiar palpus, and the short spines on the legs are very different from any other genus.
Hentz, in his descriptions of Clubiona saltabunda and C. albens, questions the generic position. The exaggerated




================================================================================

length of the legs in both sexes, the difference in the male palpi and the number and arrangement of spines on the legs do not agree with any of the genera from Central America, so the new genus Anyphsenella has been made for them. .
1. Ventral furrow one-third nearer epigastric plate than to spinnerets ............................................................ Aysha. Ventral furrow one-third nearer spinnerets than to epigastric plate, anterior median eyes largest Marcellina
Ventral furrow about midway between epigastric plate and spinnerets, anterior eyes equal or median slightly smaller ...................................................................... 2. 2. Anterior tibiae as long as both cephalothorax and abdomen ........................................................ Anyphsenella Anterior tibiae normal .................................... Anyphsena. Marcellina gen. n.
Type : Clubiona piseatoria, Hentz.
Cephalothorax oval, thoracic groove rather long, front broad and obtuse. Anterior row of eyes slightly recurved, equidistant, median largest of the eight; posterior row pro- curved and equal; quadrangle higher than wide, wider in front; lateral eyes separated by less than diameter of an- terior laterals. Clypeus not as wide as diameter of anterior median eye. First leg longer than fourth; anterior legs with few spines on ventral side. Mandibles of male well- developed with two widely separated teeth on superior margin of groove and two teeth on inferior margin. Ven- tral furrow one-third nearer spinnerets than to epigastric plate.
The genus differs from Gayenna, which has the ventral furrow near the spinnerets, by the large median eyes and from the other American genera by arrangement of the eyes, number of teeth on the mandibles and the shorter spines on the legs.




================================================================================

Psyche [ June-September
Marcellina piscatoria (Hentz)
Clubiona piscatoria Hentz, Journ. Boston Soc. Nat. Hist. ; 1847, 5, p. 450, pi. 23, f. 15 ; " 8 Alabama" ; reprint, 1875, p. 84, pi. 10, f. 15, pi. 18, f. 29.
8 7 mm. long, cephalothorax 3 mm. I leg, 15 mm. long. Cephalothorax pale yellow without markings, covered with many fine, black hairs; abdomen pale yellow without markings, with many long, fine hairs; legs pale, covered with dark hairs; spines on all legs less than diameter of joint, few and becoming irregular; more spines on upper side of legs than on ventral. Spines, I leg, tibia 2, near middle, 1 lateral, all less than diameter of joint; metatarsus 2-2, 1 lateral. Metatarsus as long as tibia. Eyes; anterior row recurved, equidistant, A. M. E. larg- est ; posterior row procurved so that lateral eyes are almost touching, equal in size; P. M. E. slightly nearer P. L. E. than to each other and smaller than A. M. E. Clypeus about as wide as diameter of A. L. E.
Mandibles, brown, porrect, one-half the length of cephalo- thorax, covered with long, black hairs. Two isolated teeth on superior margin and two isolated teeth on inferior mar- gin. Cephalothorax widest between I1 and I11 leg and only slightly narrowed in front of I coxae; thoracic groove rather long, sternum triangular, widest between coxae I and 11; coxae IV separated by less than half a diameter. Ventral furrow one-third nearer the spinnerets than to the epigastric plate.
Palpus. Femur two-thirds as long as cephalothorax, curved, tibia but little longer than patella, with apophysis near tip, as in figure 1, tarsus longer than femur and pro- longed far beyond the palpal organ; embolus prolonged in a fine straight point the length of the tarsus. 2 8 Ala.; Auburn, C. F. Baker coll.; 1 8 im.; Mass.; Newton 17 Oct., 1904, E. B. Bryant coll.



================================================================================

Anyphaena
Type : Aranea accentuata Walckenaer, 1802. Cephalothorax oval, thoracic groove rather long, front broad and obtuse. Anterior row of eyes straight or weakly recurved, the median usually the smallest; posterior row weakly procurved, equal and usually equidistant; quad- rangle of median eyes higher than wide and narrower in front; lateral eyes well separated. Legs long with scopulse on all tarsi and often on the anterior metatarsi. Ventral furrow about midway between epigastric plate and spin- nerets.
The genus Anyphsena was made by Sundevall in 1833. (Conspectus Arachnidum, p. 20) for Aranea accentuata Walckenaer. In this species the ventral furrow is a little nearer the epigastric plate than to the spinnerets and there is one pair of spines (basal) on the anterior metatarsi. The A. M. E. are smaller than the A. L. E., the mandibles are vertical, with two teeth on the inferior margin of the groove and the palpus is quite unlike any of the American forms. In a careful examination of one of the common species (A. celer Hentz), it is found that the position of the ventral furrow often varies from the middle to just in front, or a little behind the middle, so that it is not easy to use as a primary character, but in all species examined the position of the furrow is approximately the same in the two sexes, also the number and arrangement of the spines on the an- terior legs. With the exception of A. char sp. n. from Florida and A. aperta Banks from Washington all the American species have two pairs of spines 011 the first and second metatarsi. The first species also differs in having a large club-shaped process on the patella of the male palpus, a character found on at least one European species. In 1913 Berland, Araignees de l'Equateur, called atten- tion to the subdivision made by M. Simon in 1903. The first group has two teeth on the inferior margin of the mandibles and the ventral furrow is nearer the spinnerets than to the epigastric plate. The second group, which in- cludes Anyphsena, has several teeth on the inferior margin



================================================================================

106 Psyche [ June-September
(from 4 to lo), and the ventral furrow is nearer the epi- gastric plate than to the spinerets.
This character of teeth
is found variable, as in A. laticeps sp. n. there are no teeth on the superior margin of the male and a distinct carina on the inferior margin; and in the female, a carina on the superior margin and no teeth on the inferior margin, yet the male palpus is nearer the type of the genus than any other American species.
With the material available, the genus Gayenna is re- stricted to species from South America, with the ventral furrow near the spinnerets (at least one-third nearer than to the epigastric plate), and the genus Anyphsena, to species with the ventral furrow near the middle (midway between the epigastric plate and the spinnerets) and usually two pairs of spines beneath the anterior metatarsi. Future study may prove that the American species of Anyphsena may belong to some of the numerous neotropical genera, rather than to the European genus Anyphsena. It has not been possible to make a satisfactory key for the females of the genus. Possibly when both sexes of all species are known, some character can be recognized by which they can be separated, but at present they can be identified only by the vulva. . In both sexes the number and position of spines on the anterior legs is the same. Anyphaena
Males
1. I metatarsus 1-1 spines, head broad, mandibles genicu- late, large club-shaped spur on patella of palpus calcar sp. n.
I metatarsus 2-2 spines ........ .... .... .... .. ............... .... ... 2. 2. Head but little narrowed in front of I coxae, mandi- bles porrect, cephalothorax shining chestnut brown laticeps sp. n.
Head less than half as wide as cephalothorax between I1 and I11 legs, cephalothorax never brown ............ 3.



================================================================================

3. 11, I11 and IV coxae modified ................................ 4. ...................................................... All coxae normal 5. 4. 111 coxae with posterior spur bifid and with central process of palpus broad. ........................ .pectorosa Koch. 111 coxae with posterior spur slender and curved and with central process of palpus slender. .fraterna Banks. 5. I11 leg modified ........................................................ 6. I11 leg normal .......................................................... 9. 6. I11 tibia with 2 short, stout spines on ventral, pos- ............................................................ terior margin 7.
I11 tibia with but 1 short, stout spine on ventral pos- ............................................................ terior margin 8.
7. Dorsal section of tibial apophysis of palpus broad and ........................................................ triangular celer Hentz
Dorsal section of tibial apophysis slender and sharp maculata Banks.
8. Tip of ventral part of tibial apophysis fairly broad and bifid and upper part reduced to a small, sharp point scarcely noticeable .................... coloradensis sp. n. Tip of ventral part of tibial apophysis rather slender and notched, and the upper lobe fairly large, pointed ...............................
and triangular .crebrispina Chamb.
9. Distal apophysis of tibia ending in a simple, sharp point, tibia twice length of patella. .......... .ruens Chamb. "Distal apophysis in form of undivided chitinous lam- ina and narrowing distad, but with apex subtruncate ..........
and weakly notched at middle".
saniuana Chamb.
Anyphzna cakar sp. n.
8 4 mm. long; cephalothorax 1.7 mm.
Cephalothorax yellowish, darker in front with white hairs on head; abdomen pale yellow with many dark spots and streaks, which unite in the posterior median part to form a series of chevrons; legs yellowish with faint traces



================================================================================

108 Psyche [ June-September
of dark spots ; spines, I leg ; tibia 2-2, second pair at about middle of joint, 2 lateral; metatarsus 2, 1 lateral, scopulae very scant on anterior tarsi and metatarsi. Anterior row of eyes slightly recurved, equidistant, A. M. E. smallest; posterior row longer than anterior, P. M. E. slightly nearer P. L. E. than to each other, P. M. E. largest; quadrangle higher than distance between P. M. E. eyes. Clypeus narrow, scarcely more than diameter of A. M. E. eyes. Mandibles vertical, with many long, black .hairs ; superior margin of fang groove, a denticulate carina ; inferior margin a group of five minute teeth near median edge.
Ventral furrow midway between spinnerets and epi- gastric plate.
Palpus.
Patella and tibia of equal length, the former with a large club-shaped spur on the outer side, nearly as long as the joint; tibia with a long apophysis resting against the palpus, ending in a sharp spine; the ventral side of the tibia has a pronounced keel two-thirds the length of the joint.
The upper half of the palpal organ has a large white sack with a slender white filament at the upper end and a tube from the lower part which bends and disappears in the upper part of the organ. On the inner-side is a piece ending in a sharp point a little above the white sack, and in the opposite side is a slender piece ending in a curved hook.
Type : 3 8 Fla. ; Dunedin, 1927, W. S. Blatchley coll. This is the only American species of Anyphzna with a spur on the patella.
Anyphzna laticeps sp. n.
8 4 mm. long; cephalothorax 1.5 mm., abdomen 2.5 mm. Cephalothorax shining chestnut brown, a little darker about the eyes; abdomen greenish gray; legs little lighter in color than cephalothorax, long and slender with darker bands at middle and end of tibia. Spines, I tibia 2-2, long



================================================================================

and slender, no lateral, metatarsus 2-2, long and slender, no lateral.
Eyes : anterior row straight, equal and equidistant ; pos- terior row slightly procurved, longer than anterior row, P. M. E. little nearer P. L. E. than to each other; laterals separated by radius of P. L. E.; quadrangle as high as distance between P. M. E. Clypeus a little wider than diam- eter of anterior eyes:
Mandibles dark, porrect and more than half as long as cephalothorax, no teeth on superior margin of groove, dis- tinct denticulate carina on inferior margin ; fang long and sinuate; labium longer than wide, half as long as rnaxillz, and deeply notched.
Cephalothorax, narrowed but little in front of I coxae. Sternum twice as long as broad, widest between 11 coxae. Ventral furrow a little nearer epigastric plate than to spin- nerets.
Palpus.
Tibia twice as long as patella, apophysis one- third length of tibia and pressed close to tarsus. Palpal organ nearly fills the cavity.
Q 4.2 mm. long.
Cephalothorax shining chestnut brown, darker about the eyes and with scattered white hairs on sides; abdomen pale yellow with irregular dark marks arranged to form broken chevrons; venter pale; legs lighter in color than cephalothorax without the dark bands and spines as in male. Eyes as in male. Mandibles dark brown and geniculate; carina on superior margin of groove and no teeth on inferior margin. Cephalothorax narrowed but little in front of I coxae. Sternum light and shining, nearly twice as long as broad. Ventral furrow a little nearer epigastric plate than to spinnerets. Epigynum dark with two broad open- ings below two pairs of dark spots.
Type: 8 Q Ga. ; Thompson's Mills, H. Allard coll. Paratypes. 1 Q Va. ; Falls Church, N. Banks coll. ; I 2 Ala. ; Auburn, C. F. Baker coll.
The dark color of the cephalothorax would seem to ally this species to A. striata Becker, but the description of



================================================================================

110 Psyche [ June-September
that species would indicate a much larger spider, possibly an Aysha.
The two following species are separated in the males by the modified posterior coxae.
Anyphzna pectorosa L. Koch
, Die Arach. fam. Drassiden, 1866? p. 198, p1. 8, f. 131, "I 8 Baltimore."
Anyphgna calcarata Emerton, Trans. Conn. Acad. Sci. ; 1890, 8, p. 189? p. 6, f. 3, " 8
Q Conn.; West Haven, July
. . . . N. Pike's Long Island Collection.?' The IV coxae in the males have a small? pointed process directed outward; the I11 coxae have two processes, the posterior one is pointed inward and distinctly bifid? the an- terior process is a blunt tooth.
The principal process in
the palpus is broad and flattened. On the upper side of the tibia is a small spine.
This species is found from Massachusetts south to North Carolina and Tennessee and west to Missouri. Anyphzna fraterna (Banks)
Gayenna fraterna Banks, Trans. Amer. Ent. Soc.; 1896, 23? p. 63. " 8 near Sea Cliff, N. Y.
June."
Anyphmm conspema Keyserling, Verh. Zool. bot. Ges. Wien 1887, 37, p. 453, pl. 6, f. 23 " Q
Ky.; Bee Spring,"
preoccupied by sir no^^ 1878.
This species is distinctly spotted.
In the male the pos-
terior process on the I11 coxae is slender and simple, not bifid as in A. pectorosa. In the palpus the largest process is slender and curved only at the tip. The tibia has a small cone-shaped process on the inner basal side as well as a short spine on the upper. The epigynum is long and narrow instead of broad as in pectorosa. Type.
2 8 N. Y.; Sea Cliff, N. Banks Coll. Found from Massachusetts to Tennessee and Kentucky.



================================================================================

The four following species form a natural group, with the third leg in the male modified, with the third femur en- larged, and one or two stout, cone-shaped spines on the posterior ventral side of the tibia.
In both sexes there is
a distinct dark ring near the distal end of each femur. Anyphzena celer ( Hentz )
Figs. 7, 25, 28
CZubiona celer Hentz? Journ. Boston Soc. Nat. Hist.; 1847, 5, p. 45z7 pl.,. 23, f. 20; reprint, p. 87, pl. 10, f. 20. "North Carolina, Alabama."
Anyphzna i~xcerta Keyserling, Verh. zool. bot. Ges. Wien, 1887, 37, p. 452, pl. 6, f. 22. " 9 Mass. ; Cambridge.'' A widely distributed species found under dead leaves from New England to Texas. The position of the ventral furrow varies from mid-way between the spinnerets and the epigastric plate and nearer to the epigastric plate. The under side of the tibia of the third', leg has two very short spines on the posterior side.
A common species from Massachusetts south to Texas. Anyphama maculata (Banks)
Figs. 8, 31
Gayenm maczdata Trans. Amer. Ent. Soc.; " 8 D. C.;
Washington.'' 1896, 23, p. 64. S. C. Bishop and Crosby, Journ. Mitch. Soc. 1926, 46, p. 189, pl. 24, f. 37, 38. Crosby and Bishop give a very full description of the male, but fail to mention the enlarged third femur and the two stout spines on the tibia of the third leg. The palpus
is very similar to that of A. celer, but the tibia1 apophysis is quite distinct. The upper branch is shorter and ends in a sharp point, and the inner branch is a thin leaf-like plate, folded almost double. The large rounded protuberance on the basal half is larger and more conspicuous than in A. ceZer.
Type: 8- D. C. Washington.




================================================================================

112 Psyche [ June-September
8 s N. Y. ; Sea Cliff, N. Banks Coll.; s Va.; Falls Church, N. Banks Coll. 1 Ky.; Hart Co.; Rio. A. F. Archer coll.
Anyphaena coloradensis sp. n.
s 4.5 mm. long.
Cephalothorax light brown with faint dark marks from the lateral eyes to the posterior margin; abdomen light yellow with five median dark bars slightly bent as chevrons, sides and venter with dark spots and streaks; sternum light; legs light with dark spots at base of spines; spines I leg, tibia, 2-2-2 none apical, 2 lateral, metatarsus, 2-2, 2 lateral, metatarsus shorter than tibia; coxae I11 and IV densely, minutely, spinulate, femur I11 enlarged and on ventral surface, minutely spinulate; two long spines on dorsal surface and one very short, stout spine on posterior, tibia I11 two long spines on anterior edge and one stout spine posteriorly, two lateral.
Ventral furrow is nearer
spinnerets than to epigastric plate. Eyes; anterior row re- curved, A. M. E. smallest and 'less than their diameter apart, and less than radius from the A. L. E. ; posterior row procurved, P. M. E. nearer P. L. E. than to each other; laterals separated by less than radius of A. L. E.; quad- rangle nearly twice as high as distance between posterior median.
Palpus:
Patella as long as tibia; tibia1 apophysis pro- longed half the length of the tarsus, with upper edge ser- rate, with one distinct tooth midway, palpal organs filling the entire cavity and similar to A. celer. Q 4.7 mm. long.
Markings the same as in the male, but not quite as dis- tinct, except on the legs where the dark marks are more pronounced. The posterior coxae are not roughened or the I11 femur enlarged.
Eyes same as in the male. Epi-
gynum as figured.
Type: 1
8, 2 8 Col. ; Boulder, T. D. A. Cockerel1 coll. N. Banks Coll.




================================================================================

Anyphaena crebrispina Chamberlin
Pomona Coll. Journ. Ent. Zool.; 1919, 12, p. 10, pi. 4, f. 4. " 8 Cal. ; Claremont", No. 353.
The third tibia has but one stout spine on the posterior side. . The tibia1 apophysis has the lower branch about one- third the length of the tarsus, with a distinct notch at the tip and the upper lobe in a pointed triangle. Type :
1 8 Cal. ; Claremont, R. V. Chamberlin Coll. This is probably the male of Anyphzna californica (Banks) as the later has been found at Claremont.
Anyphsena ruens Chamberlin
Pomona Coll. Journ. Ent. Zool.; 1919, 12, p. 11, pi. 5, f. 1. " 8 Cal.; Claremont, April, 1913, No. 352." Anyphsena mundella Chamberlin, ibid, p. 12, pi. 5, f. 3. ? nee 8 "Cal. ; Claremont."
In the original description of A. mundella the female only was described and the palpus of the male figured. On examining the specimens, the two obviously belong to differ- ent genera. The male has but one pair of spines on the anterior metatarsi, a broad low cephalic region and the ven- tral furrow quite near the epigastric plate, all characters of Aysha. The female has two pairs of spines on the an- terior metatarsi, the narrow and high cephalic portion and the ventral furrow at "middle or slightly behind middle of abdomen." In all species of Anyphsena seen the number of spines on the anterior legs, shape of cephalothorax and the position of the ventral furrow is the same in both sexes. Type: 1 8 M. C. Z. no. 352, Cal.; Claremont. 1 9. M. C. Z. no. 348, Cal.; Claremont.
8 B. C.; Victoria, Taylor coll.; 8 Cal.; Claremont; C. F. Baker coll. ; Mt. Helens ; 8 July, 1918. Helen Van
Duzee coll.




================================================================================

114 Psyche C June-September
The following species are known only from females. Anyphaena aperta (Banks)
Gayenna aperta Banks, Proc. Cal. Acad. Sci.; 1921, 11, p. 100, f. 3. " Q Wash. ; Olympia."
The ventral furrow is midway between epigastric plate and spinnerets. The spines on I tibia 2-2-2 (one pair distal), 2 lateral, metatarsus 2, 2 lateral. Type : 1 s Wash. ; Olympia, T. Kincaid coll. 1 Q Wash. ; Blakley Is. ; 8 July 1928. L. G. Worley coll. Anyphaana californica (Banks)
Gayenna californica Banks, Proc. Cal. Acad. Sci.; 1904, 3, p. 338, pi. 38, f. 2.
" Q Palo Alto, Mill Valley."
Anyphm intermontana Chamberlin, Can. ent. ; 1920, 52, p. 200, f. 6.
Type : 1 Q Cal. ; Palo Alto, R. W. Doane coll. 1 Q Cal.; Marin Co.; Ross, 3 Q Claremont, N. Banks Coll. 2 Q Utah ; Mill Creek, R. V. Chamberlin coll. Anyphaana fragilis Banks
9
Can. Ent. ; 1897, 29, p. 194 " Q Fla. ; Jacksonville.'' Gayenna parvula Banks, Proc. Ent. Soc. Wash.; 1899, 4, p. 191. " Q La. ; Shreveport."
Cephalothorax brown, darker about the eyes with scat- tered white hairs, cephalic groove short; abdomen pale, sometimes with a few marks bent almost to form chevrons posteriorly; venter pale without marks; legs pale, some- times with traces of two dark rings on anterior tibiae, and at end of metatarsi.
Spines; I leg, tibia, 2-2, long, no lat- eral metatarsus, 2-2, long, no lateral. Cephalothorax very broad at anterior margin, with only a slight narrowing in front of coxae I. Mandibles geniculate and very dark brown, covered short hairs.




================================================================================

Eyes; anterior row straight, equidistant and A. M. E. a little smaller than A. L. E. ; posterior row procurved, equal and P. M. E. a little nearer to P. L. E. than to each other. Quadrangle but little higher than wide and narrowed in front.
Clypeus low, but little wider than diameter of A. M. E.
Ventral furrow a little nearer epigastric plate than to spinnerets.
Type : 2 $ , 1 8 im. Fla. ; Jacksonville ; 2 $ La. ; Shreve- port. N. Banks Coll. ; 1 ? Ga. ; Bainbridge, J. C. Bradley coll.
Anyphaena pacifica (Banks)
Gayenna pacifica Banks, Trans. Amer. Ent. Soc.; 1896, 23, p. 63. " $ Wash.; Olympia."
Type :
3
$ , 1 8 im. Wash. ; Olympia, J. Kincaid coll. ; 1 $
Idaho; Moscow Mts., 29 Oct. 1910. J. A. Hyslop coll.; 2 Q Wash.; Wawawai, W. M. Mann coll.; 1 $ Cal.; Mt. Tamalpais, Gal. Acad. Sci. Coll.
Anyphaena zina Chamberlin
Pomona Coll. Journ. Ent. Zool.; 1919, 12, p. 11, pi. 4, f. 5. Type : 1 9 Cal. ; Claremont, R. V. Chamberlin coll. Anyphamella gen. n.
Type : Clubiona saltabunda Hentz.
Eyes similar to Anyphaena; A. M. E. smaller than A. L. E.
Mandibles vertical on both sexes, with row of many minute teeth on the inferior margin, and a row of four dis- tinct teeth on the superior margin. First leg longest and tibia longer than or nearly as long as the entire length of spider. Two pairs of spines beneath I tibia and two pairs beneath I metatarsus. Ventral furrow about midway be- tween spinnerets and epigastric plate.
It differs from Wulfila in the number of spines on the metatarsi and from Sillus in the much longer legs and the structure of the male palpi.




================================================================================

Psyche [ June-September
Anyphaenella saltabunda (Hentz)
Clubiona ? saltabunda Hentz, Journ. Boston Soc. Nat. Hist.; 1847, 5, p. 453, pi. 23, f. 23; "Alabama". reprint, p. 89, pi. 10, f. 23, pi. 18, f. 19.
Anyphs~na saltabunda Emerton, Trans. Conn. Acad. ; 1890, 8, p. 187, pi. 6. f. 4.
s 3.5 mm. long; 9 4 mm. long.
Cephalothorax, abdomen and legs light yellow with gray, broken stripes on cephalothorax, and a broken marginal band; two parallel rows of gray dots on the abdomen. Ster- num light with a scalloped dark mark around edge. , Legs very long and slender.
I metatarsus as long as
I tibia.
Spines, I tibia, 2-2-2. 2 lateral; metatarsus, 2-2, 3 lateral. Front less than half the width of cephalothorax between I11 legs. Eyes; anterior row straight, A. M. E. smallest; posterior row procurved, equal, P. M. E. nearer P. L. E. than to each other; lateral eyes almost touching. Clypeus width of A. M. E. Mandibles small and vertical, with four teeth on superior margin and with many, minute ones on the inferior. Labium slightly notched. Ventral furrow midway between epigastric plate and spinnerets. Palpus:
tibia more than twice as long as patella, much bent and the apophysis near the base.
Epigastric plate extends nearly across the abdomen. Found from Massachusetts to Florida and west to Wyoming.
Anyphaenella alba (Hentz)
Clubiona ? albens Hentz. Journ. Boston Soc. Nat. Hist.; 1847, 5, p. 454,' pi. 23, f. 24; "Alabama" reprint, p. 89, pi. 10,f. 24, pl. 18, f. 32.
Chiracanthium albens Hentz, Banks, Proc. Phila. Acad. ; 1904, p. 123.
s 4.5 mm. long ; $ 4 mm.
Cephalothorax, abdomen and legs almost colorless and without markings. Legs very long and slender, I metatar-



================================================================================

sus longer than I tibia. Spines, I tibia, 2-2, 2 lateral, meta- tarsus 2-2, 2 lateral. Eyes ; anterior row slightly recurved, A. M. E. smallest and less than a diameter apart, and nearer to each other than to the A. L. E.; posterior row, procurved, equal and equidistant, oval and surrounded by black, quadrangle much higher than wide, anterior side about half as wide as posterior. Mandibles small and verti- cal.
Ventral furrow a little nearer epigastric plate than to spinnerets.
8 D. C. ; Washington, N. Banks Coll. ; Va. ; Falls Church, N. Banks Coll.; N. C. Raleigh, C. S. Brimley coll. Aug. 1916. 1
Ky. ; Bee Spring, F. G. Sanborn; 1 s Fla. ; Sebastian, 17 Apr. 1919, G. Nelson coll.
Aysha
Type : A ysha prospera Keyserling.
Cephalothorax low and broad, only slightly narrowed in front of the first coxae. Posterior row of eyes almost straight, anterior row subequal; clypeus about as wide as diameter of anterior median eye. First leg usually longer than fourth, one pair of spines beneath on first metatarsi. Ventral furrow near epigastric plate, at least one-third nearer than to spinnerets. The male palpus has the em- bolus in a sweeping circular curve, and the epigynum with a distinct circular chitinous rim.
F. 0. P. Cambridge states (Biol. Cent. Amer., 1900, 2, p. 98), that the lower fang groove of the mandible has 3-7 teeth. In A. cambridgei sp. n. there are no teeth on the lower groove, and the upper groove has a denticulate carina, instead of the usual three or four distinct teeth. Males
1. Tibia of palpus with 1 spur ........................................ 2 Tibia of palpus with 2 spurs ...................................... 5 2. Tibia much longer than tarsus, spur bifid at tip longipalpus sp. n.
Tibia plainly not longer than tarsus ........................ 3




================================================================================

118 Psyche June-September
3.
Tibia shorter than tarsus, spur as long as tibia. I tibia 2-2. 1 lateral, metatarsus 2, 1 lateral.. . .g~acilis Hentz Tibia about as long as tarsus, spur shorter than tibia 4
4. I leg, tibia 2-2-2. 2 lat., metatarsus 2, 1 lateral. Spur with large hook at tip ................................. .velox Becker. I leg, tibia, 2-2, no lat., metatarsus 1-1, no. lat., spur ........
sinuous with minute hook at tip
cambridgei sp. n.
5.
Upper spur as long as tibia and with rest of joint as long as tarsus.. ......................................... .minuta Camb. Upper spur shorter than tibia but with tibia longer than tarsus.. ............................................... .decepta Banks Aysha longipalpus sp. n.
8 10 mm. long. ceph. 4 mm.
Cephalothorax light brown with scattering white hairs; abdomen light grey without markings; legs light brown, first and second darkest. Spines, I, leg, tibia, 2-2-2, (one pair distal), no lateral; metatarsus, 2, no lateral. I femur 5 mm., patella 1.5 mm. ; tibia 7.5 mm., metatarsus 8 mm., tarsus 2 mm. Eyes; anterior row slightly recurved, equal and equidistant; posterior row procurved, equal and equi- distant; lateral eyes radius of P. L. E. apart. Clypeus slightly more than radius of anterior eyes. Mandibles por- rect, two-thirds as long as cephalothorax; four teeth on superior margin and a row of minute teeth on inferior mar- gin; fang long; sternum widest between II and I11 coxae and prolonged between the IV coxae. Ventral furrow very near epigastric plate.
Palpus.
Tibia much longer than tarsus and more than twice as long as patella; very slender, apophysis bifid at tip and little longer than diameter of the segment. Ap- pendages of the palpal organ confined to the upper half of the bulb.
Type : 1 8 Costa Rica; San Jose, Valerio coll. Differs from A. tennis (L. Koch) Fig. 19, by the longer and more slender tibia, notched tibia1 apophysis and smaller palpal organ.




================================================================================

Aysha gracilis (Hentz)
Clubiona gracilis Hentz, Journ. Boston Soc. Nat. Hist. ; 1847, 5, p. 452, pi. 23, f. 19; reprint, p. 86, pi. 10, f. 19, pi. 18, f. 8. "North Carolina, Alabama." Anyphma rubra Emerton, Trans. Conn. Acad., 1890, 8, p. 186, pi. 6, f. 1. " 9, Massachusetts and Connecticut and in Pike's Long Island Collection." ibid, 1909, 14, p. 220, pi. 9, f. 8.
A ysha orlandensis Tullgren.
Bih. Svensk. Akad. ; 1901,
27, p. 19. f. 4. " 9 Fla. ; Orlando."
A widely distributed species from Massachusetts to Flor- ida and as far west as Nebraska.
Aysha velox (Becker)
Anyphma velox Becker, Ann. Soc. Ent. Belg. ; 1879, 22, p. 83, pi. 2, f. 5-7. $ "Mississippi; near Pascagoula." Anyphsena floridana Banks, Trans. Amer. Ent. Soc. ; 1896, 23, p. 63. " 9 Fla.; Lake Worth."
Chiracanthium falcdum Chamberlin, Bull. Mus. Comp. ~ool.'; 1925, 67, p. 220. " a: Fla. ; Sebastian." The type, a female, is from Miss. It is the common Aysha in Florida, Alabama and Louisiana, and is found in Bermuda, Cuba and was found by Mr. Banks on Taboga Island, Panama, 29 June, 1925.
Aysha cambridgei sp. n.
a: 8.5 mm. long; ceph. 3 mm.
Cephalothorax light brown ; abdomen yellowish grey without markings ; legs light brown, lighter than cephalo-



================================================================================

120 Psyche [ June-September
thorax. Spines, I leg, tibia 2-2, no lateral, metatarsus 2, no lateral. I femur 4 mm., patella 1.8 mm., tibia 4.8 mm. ; metatarsus 4 mm., tarsus 2 mm. Eyes; anterior row slightly recurved, equal and equidistant, posterior row pro- curved, equal and equidistant, lateral eyes separated by diameter of P. L. E. Posterior eyes not surrounded by black. Mandibles brown, less than one-half as long as cephalothorax ; porrect ; denticulate carina on superior mar- gin and teeth and carina lacking on inferior margin. Maxillae much broadened above the emargination, labium notched at tip. Sternum twice as long as broad. Palpus. Tibia not as long as tarsus, Apophysis two- thirds as long as tibia ending in a small hook. Appendages
of palpal organ filling three-quarters of upper portion of tarsus.
Type : 1 8 Mexico ; Guanajuato. N. Banks coll. The palpus is similar to A. velox Becker, but differs in the larger and heavier tibia1 apophysis. The spines on the
anterior legs are quite different, also the teeth on the man- dibles.
Aysha minuta F. 0. P. Cambridge
Biol. Cent. Amer. 1900, 2, p. 99, pi. 7, f. 18, 19. M. C. Z. Coll. 1 8 Mex. ; Mexico City. 1 $ Costa Rica, San Jose. Tristan Coll.
Aysha decepta (Banks)
Anyphasna decepta Banks, Proc. ent. soc. Wash.; 1899, 4, p. 190. " $ La., Shreveport; Texas, Brazos Co." Anyphasna incursa Chamberlin, Pomona Coll. Journ. Ent. Zool.; 1919, 12, p. 12, pi. 5, f. 2. " $ Cal. ; Clare- mont."
Anyphasna mundella Chamberlin, ibid, p. 12, pi. 5, f. 4. 8 nee. $ . "Cal. ; Claremont."




================================================================================

Anyphsena johnstoni Chamberlin, Proc. Cal. Acad., 1924. 12, p. 662, f. 105, 106.
Gulf of California.
Found in Louisiana, Texas and California. This species may prove to be Aysha simplex Cambridge, Biol. Cent. Amer., 1897, 1, p. 227, pi. 29, f. 4, 5. Panama, Veraguas. Cambridge says, "Palpi, short, slender, cubital joint short, not half the length of the radial, which is very lightly curved, cylindrical, furnished with a few long, slender spine-like bristles, and has a tapering somewhat sinuous reddish-yellow brown apophysis projecting nearly at right angles to the joint at its fore extremity on the outer side. This apophysis is longer than the width of the joint." In the Spiders of Porto Rico, Trans. Conn. Acad. ; 1930, 31, p. 61, Mr. Petrunkevitch has placed Aysha simplex Cambridge as a synonym of Aysha tenuis L. Koch. From
the number of specimens in the Museum of Comparative Zoology collection from various islands in the West Indies, the tibia of the male palpus of Aysha tenuis is longer than the tarsus, and at least five times as long as the diameter; there is but one tibia1 apophysis, not two, which is com- paratively short, the embolus occupies the upper half of the palpal organ.
From the two figures of Aysha simplex Cambridge, the tarsus is longer than the tibia, which is about three times the diameter; there are two apophyses and the embolus occupies more than half the palpal organ. It differs from Aysha decepta (Banks) by the shorter upper apophysis and the relatively smaller palpal organ. Aysha nigrifrons (Chamberlain and Woodbury) Anyphsena nigrifrons Chamberlain and Woodbury, Proc. Biol. Soc. Wash.; 1929, 42, p. 137, pi. 1, f. 4. "Utah,
St. George, Dec. 1925."
The figure of the epigynum, number of teeth on the inferior margin of the mandibles and number of spines on the anterior metatarsi show that this species is an Aysha rather than Anyphsena.




================================================================================

Psyche
PLATE 6
Fig. 1. Marcellina piscatoria (Hentz) lateral view of left palpus.
Fig.
2. Marcellma piscatoria (Hentz) dorsal view of eyes. Fig. 3. Anyphseneb calcar sp. n. ventral view of palpus. Fig. 4. Anyphsena laticeps sp. n. ventral view of palpus. Fig. 5. Anyphsena pectorosa L. Koch lateral view of palpus. Fig. 6. Anyphasna fraterna (Banks) lateral view of palpus. Fig. 7. Anyphsena celer (Hentz) lateral view of palpus. Fig. 8. Anyphsena maculata (Banks) lateral view of palpus.
Fig. 9. Anyphsena coloradensis sp.n. lateral view of palpus. Fig. 10. Anyphsena coloradensis sp. n. ventral view of palpus.
Fig. 11. Anyphsena crebispina Chamberlin lateral view of palpus.
Fig. 12. Aysha
Fig. 13. Aysha
Fig. 14. A ysha
Fig. 15. Aysha
Fig. 16. Aysha
Fig. 17. Aysha
PLATE 7
longipalpus sp. n. lateral view of palpus. gracilis (Hentz) lateral view of palpus. velow (Becker) lateral view of palpus.
cambridgei sp. n. lateral view of palpus. decepta (Banks) lateral view of palpus.
minuta Cambridge lateral view of palpus. Fig. 18. Anyphsenella saltubunda (Hentz) lateral view of palpus.
Fig. 19. Aysha tenuis (L. Koch) lateral view of palpus. Fig. 20. Anyphsenella alba (Hentz) epigynum.



================================================================================

Fig. 21. Anyphssnella alba (Hentz) lateral view of palpus. Fig. 22. Anyphsenella saltabunda (Hentz) epigynum.
Fig. 23. Anyphasna fraterna (Banks) epigynum. Fig. 24. Anyphsena laticeps sp. n. epigynum. Fig. 25. Anyphsena celer (Hentz) third left leg. Fig. 26. Aysha gracilis (Hentz) epigynum. Fig. 27. Aysha decepta (Banks) epigynum. Fig. 28. Anyphsena celer (Hentz) epigynum. Fig. 29. Anyphsena californica (Banks) epigynum. Fig. 30. Anyphma coloradensis sp. n. epigynum. Fig. 31. Anyphsena rnaculata (Banks) epigynum. Fig. 32. Anyphsena fragilis Banks epigynum. Fig. 33. Anyphsena coloradensis sp. n. third left leg. Fig. 34. Aysha velox (Becker) epigynum.
Fig. 35. Anyphsena aperta (Banks) epigynum. Fig. 36. Anyphsena pacifica (Banks) epigynum.



================================================================================

124
Psyche, 1931
Psyche [ June-September
VOL. 38, PLATE 6
Bryant, Anyphaeninae




================================================================================

19311 Anyphseninss 125
Psyche, 1931
VOL. 38, PLATE 7
Bryant, Anyphaeninae




================================================================================

126
Psyche, 1931
Psyche
[ June-September
VOL. 38, PLATE 8
Bryant, Anyphseninse




================================================================================