P. J. Darlington, Jr.
West Indian Carabidae II.: Itinerary of 1934; Forests of Haiti; New Species; and A New Key to Colpodes.
Psyche 42(4):167-215, 1935.

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PSYCHE
VOL. XLII DECEMBER, 1935 No. 4
WEST INDIAN CARABIDE 11. : ITINERARY OF 1934 ; FORESTS OF HAITI; NEW SPECIES; AND
A NEW KEY TO COLPODES*
BY P. J. DARLINGTON, JR.
Museum of Comparative Zoology
In this, my second paper on West Indian Carabidse,' are presented the results of a collecting trip to Cuba, Jamaica, and Haiti made between July 28 and November 28, 1934, through a grant from the Milton Fund of Harvard Uni- versity. I hope to continue this series of papers from time to time as additional material becomes available, and eventually to end it with a complete revision of the Carabidse of the West Indies, with discussions of their distribution and relationships, of the origin of the faunae of the isolated mountain ranges, and of other problems. ITINERARY: From August 2 to 12 aided by a small addi- tional grant from the Atkins Fund, I was at the Harvard station at Soledad, near Cienfuegos, Cuba, collecting espe- cially the smaller, more inconspicuous ground insects. On August 13 I arrived in Kingston, Jamaica, and went at once to Whitfield Hall (Miss. G. L. Stedman, office in Kings- ton) at about 4,500 ft. elevation on the south slope of the main range of the Blue Mts. From here it was easy to climb to fine collecting in the damp cloud forest of Blue Mt. Forest Reserve, and to reach Blue Mt. Peak, 7,388 ft., the highest summit on the island. On August 20 I re- turned to Kingston and motored across the island to Ocho Rios, on the north coast. Swamp and pond collecting *The publication of this article has been financed by a grant from the Museum of Comparative Zoology.
The first, "New West Indian Carabidse, with a List of the Cuban Species, appeared in PSYCHE, Vol. 41, 1934, pp. 66-131.



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168 Psyche [December
proved to be good, especially at Bogue, a large coconut plantation. Castle Daly (August 24-25, as the guest of Mr. Robert W. Bell) and Moneayue Lake (August 25-26) proved, entomologically, rather disappointing. On the 26th I returned to Kingston, staying at Mona Great House, near Hope Gardens, and spent the next three days working the Liffuanea Plain and nearby country. Ground beetles were unexpectedly numerous. A trip (afternoon of August 29) to the Rio Cobre at a point about 5 miles above Spanish- town, and to the swamps beside the main road from Kings- ton to Spanishtown, resulted in an especially fine collec- tion.
On August 30 I reached Haiti, meeting Dr. Marston Bates at Port-au-Prince, where we made our headquarters at the Sans Souci Hotel. After several days of delay, spent partly in collecting in the Cul de Sac region, we drove in a 1927 Buick roadster into- northern Haiti. From Septem- ber 5 to 11 we were at Ennery, about 1,000 ft. altitude, working especially along the fine little river near the town. On September 9 we made a rather hasty side trip to Mt. Basil, probably the highest mountain in northern Haiti, and found several fine, new mountain Carabidse in patches of very wet, low cloud forest on the summit plateau, about 4,700 ft. Engine trouble forced us to return to Port-au- Prince on September 11. We broke the drive south long enough for me to spend a very profitable four hours along the edge of the extensive swamps north of Dessalines. While our car was garaged, we persuaded Mr. Andre Audant, government entomologi~t of Haiti, to join us in a miniature "safari" to the neighborhood of La Visite, on the western end of the main range of La Selle, south of Port-au-Prince. Our schedule was arranged to the last detail through the good offices of Captain Frederick Baker, whose knowledge of the country and country people is un- surpassed. We were away from September 16 to 23, mak- ing our base camp in tall pine forest at over 6,000 ft., be- side the small river called by the local Negroes the Rivikre Blanche, but referred to by Wetmore' as the Riviere 'Birds of Haiti and the Dominican Republic, by Alexander Wet- more and Bradshaw H. Swales, Bulletin 156, United States National Museum, 1931.




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1935 ] West Indian Carabidse 169
Chotard. This stream flows south, and cannot be the north-flowing Riviere Blanche of the map.3 We used Wet- more's actual camp site (1. c., Plate 9). Ground collecting in the wet cloud forest was exceedingly good, but the pine forest, under which the ground cover had been burned, was barren.
Back at Port-au-Prince, we found ourselves unable to afford further auto travel, and Dr. Bates was forced to leave for Panama. After several days of delay, during which, through the great kindness of Dr. H. D. Barker, I was able to make auto trips to about 2,000 ft. in the foot- hills of the La Selle massif above Port-au-Prince (October 2), to the Riviere Froide about 6 miles south of the city (October 3), and to Poste Terre Rouge, 2,000 ft., in the mountains just north of the Cul de Sac (October 5), I left by public bus for Aux Cayes, on the southwestern penin- sula of Haiti. From Aux Cayes I hired a car inland to Camp Perrin, which I reached October 8. Two days suf- ficed to pacify the local authorities and hire porters, and the next 16 days were spent on foot in the maze of wet ranges which constitute the Massif de la Hotte. Our route lay first north from Camp Perrin along the narrow, new road, impassable for cars in rainy weather, which crosses the foothills toward Jeremie, then west by vile foot trails, and finally north to Desbarriere (about 4,000 ft.), on a spur which curves down first north and then west from the northern slope of La Hotte itself. From Desbarriere, October 13, with a local man as guide, I ascended the ridge to above Roche Croix (a prominent rock marked with a rude natural cross), to about 5,000 ft., and found, beside new beetles, a new genus of snake and a new lizard of a genus previously known only from Jamaica. Further ad- vance by way of the long ridge proved impracticable-the ground was covered with treacherous limestone forma- tions hidden under deep moss and bracken-so we packed and moved from Desbarriere down a thousand feet or so to Tardieu (pronounced more like "Targi" by the inhabi- tants) in the valley of the turbulent Riviere Tardieu, be- ^We used the Carte de la Republique d'Haiti, issued by the Direc- tion Generate des Travaux Publics, Port-au-Prince, 1928.



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170 Psyche [December
tween the ridge of Desbarriere and the north slope of La Hotte. From Tardieu I ascended the main peak (Pic de Macaya on the map, but called merely "La Hotte" locally; 7,800 ft. or a little less), taking four days (October 15-18) for the climb and descent. I spent an exceedingly cold night on top, with no cover except a leather jacket, and encountered various difficulties of no particular scientific interest. The climax of the ascent was partly spoiled by the discovery that surveyors had been up two or three years before, cutting a temporary trail from the other side, and had felled an acre or two of the big pines on the highest point to make room for surveying targets, but my catch of ground insects was more than satisfactory. From Oc- tober 19 to 22 our departure from Tardieu was blocked by the river, flooded by heavy rains; then two days of steady walking took us back to Camp Perrin. On October 26 and again on the 27th I collected along the shores of Etang Lachaux, a fine, small lake an hour's walk over a ridge east of Camp Perrin. This was perhaps the best single locality I found below 1,000 ft. for ground collecting. Mira- goane (October 30 to November 2), on the other hand, proved to be about the worst, in spite of the extensive swamps along the lake shore. On November 2 I reached Port-au-Prince again. Four days later an attack of ma- laria forced me to go up to Kenskoff, above the city, on one of the outer ridges of the Massif de la Selle, at about 4,500 ft., to rest and recuperate in the cool climate. I did a little collecting in the neighborhood, from about 4,000 ft. to Morne Tranchant, about 6,000, directly above Kens- koff. From November 15 to 20, after returning to Port- au-Prince, I made a very unprofitable trip to Manneville, at the western end of Etang Saumiitre, and to swampy Trou Caiman nearby, and an equally unprofitable ascent of Mt. Trou d'Eau
(about 5,200 ft.). This is the highest
Haitian peak just north of the Cul de Sac, near Thoma- zeau. On November 22 I sailed from Port-au-Prince for New York.
PRESENT FORESTS OF HAITI: Four centuries of occupation by a large population of Negro peasants (according to the World Almanac there are about 2,550,000 people in the country to-day, or about 250 per square mile) have left no



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1935) West Indian Carabidas 171
real forest at low altitude in Haiti, nothing better than cactus and acacia scrub and dry, open woodland, and even these exist only where fresh water is not available for humans. A single exception should perhaps be made of the tip of the southwestern (Tiberon) peninsula, near Dame Marie. Here, according to Dr. Barker, who has seen the region from the air, good forest seems to be continuous from the westernmost ranges of the Massif de la Hotte down almost to sea level. The vegetation indicates a local rainfall of nearly 200 inches a, year. With this exception, all the wet forest of Haiti (I am speaking of the country, not the island, for I do not know Santo Domingo) is now confined to the mountains.
The mountains of the country of Haiti form three nab ural geographical divisions. Each division has its own distinctive ground fauna, found as a rule only in the rem- nants of cloud forest at high altitudes. Almost all of the true mountain species and even some of the genera are restricted to a single one of these divisions. This is true not only of the insects but of the lizards, frogs, mollusks, Peripatus, etc. as well. The mountains of the northern part of Haiti, north of the Cul de Sac, may probably be considered faunistically as a single division (Division I.). They are relatively low and accessible, and what little forest is left on them is confined to the summits of a very few ranges. The sooner these mountains are thoroughly explored zoologically, the better, for some forest species have probably already become extinct, and others will fol- low. On the plateau of Mt. Basil (4,700 ft), to which we climbed from the road between Ennery and St. Michel de I'Atalaye, there is still a good deal of low, dense, wet cloud forest. The plateau is rough and there is no permanent drinking water, and the Negroes do not like the cold at that altitude, but even so they climb up daily and are grad- ually clearing the best growth to plant vegetables. In 1928 there were some patches of much better cloud forest on Haut Piton (about 3,900 ft.) , near Port-de-Paix, according to Mr. James Bond, who climbed the mountain on an or- nithological reconnaissance, but they were rapidly being destroyed at that time. There is also a small area of forest on Fuilboreau, just north of Emery, but it is said to be



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172 Psyche [December
on a slope, and much drier than the plateau forests. I have .
not been able to learn definitely of any other good forest in the North, although a little may still exist on other iso- lated ranges. The summit of Morne Salnave (NW. Haiti) is said to be covered with low, scrubby second growth (Bond). The conditions I found on Mt. Trou d'Eau (5,200 ft.) are probably typical of most of the more accessible peaks: very dry, rather open woods on the lower slopes; pastures and gardens above, where the soil is better, xig'ht up to the summit, which was crowned by a corn field. South of the Cul de Sac, which forms a perfect barrier to mountain animals, the mountains are higher and wet- ter, and there are still large areas of forest. This is the case on the Massif de la Selle (Division II.) , or at least on the plateau of the main range, where there are extensive pine woods as well as long strips of dense, low cloud forest. The cloud forest, under which the soil is black and rich, ia being attacked by Negro farmers, who have pushed their gardens in places up to 7,000 ft,, but it will probably be many years before it is all destroyed. The pine woods "will probably last even longer, for the soil is poor, but unfor- tunately, at least on the western end of the range, which was all we saw, they have been marred by repeated burn- ing of the ground vegetation under the trees. Some of the outlying ridges, too, of this massif, have evidently sup- ported both pine and cloud forest in the past, but for the most part, as on Morne Tranchant above Kenskoff, the trees have all been cut and the mountain fauna persists, if at all, only in damp thickets and gullies. There is, how- ever, some good cloud forest left in the Crete Piquant (western) section of the La SeIIe massif (Bond). West of the higher ranges of the Massif dc la Selle, for perhaps 60 miles along the narrow middle part of the peninsula, lie a series of lower ridges without notable peaks. These, as can be seen from the road, are mostly either stripped or so dry as never to have been heavily forested. The latter is probably the case, for the ground life of the La Hotte mountain complex (Division III.) at the outer end of the peninsula has evidently long been isolated from that of La Selle. First of the important western moun- tains is Bonnet Carre, between Aquin and L'Asile, a broad



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19351 West Indian Carabidse 173
dome a little over 4,000 ft. high. Seen from La Hotte, this mountain seems still to be capped with at least a square mile of dark forest. It is entirely unknown entomological- ly. Botanically it is related to La Hotte, but with some peculiar forms. Between Bonnet Carre and the eastern- most rainforest (at about 2,000 ft.) on La Hotte there are 20 miles or more of low, dry ridges. The high ranges of the central part of the La Hotte massif (Pic de Macaya, Pic du Formon, and Pic de la Gde. Colline on the map), and apparently also most of the lower ranges to the west almost to the coast, are very heavily forested. On La Hotte itself is a completely unbroken tract, without even trails, probably 8 or 10 miles across the narrowest way. The forest is of two kinds: fine pine with unusually thick un- dergrowth and with thick moss and pine needles on the ground, and lower, denser, very wet cloud forest, without pine, fringed with cutting climbing bamboo and other atrocious plants. It is in the La Hotte region, of all Haiti, that there is to-day the best chance of finding novel forms of life, and it is undoubtedly there that natural conditions will persist longest.
DESCRIPTIONS : Wherever in the following pages definite proportions are given-for instance the relative width of head and prothorax-they are based on actual measure- ments made under a binocular microscope. Estimates made without measuring are likely to be surprisingly inaccurate. I have invariably measured the width of the head across the eyes at the widest point; width of prothorax, at widest point; length of prothorax, at middle, regardless of whether or not the anterior angles project forward; width of base of prothorax, between angles ; width of apex, between most advanced points of angles.
Ardis tomus alti~cola n. sp.
Very stout and convex; black, shining but not metallic; not spotted; legs piceous ; antennae, palpi, and tarsi rufous. Head with front margin of clypeus faintly convex; front lightly, vertex not distinctly transversely impressed. Pro- thorax suborbicular, moderately narrowed in front; lateral margins distinct to base; disk with usual fine middle line and anterior transverse impression ; not punctate. Elytra



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174 Psyche [December
broad and convex; humeri rounded; striae deep, impunc- tate, entire and strong at apex, 2nd as well as others entire at base; short accessory stria at extreme base close to su- ture; 3rd stria running to humeral margin; intervals con- vex, shining, 3rd with 5 setigerous punctures; trace of alutaceous microsculpture at extreme base of elytra. Last ventral with 2 setigerous punctures each side near margin. Front tibia with 1 strong and 1 weak tooth on outer side above terminal digit; front tarsi rather widely dilated. Length 4.7; width 1.8 mm.
HAITI: holotype (M. C. Z. no. 22012) and 1 paratype from Mt. Bourette, La Selle massif, 5,000 ft., Sept. 16 & 23; under mossy stones in a damp gully.
Very similar to A. Isevistriatus F. & S. of Guadeloupe (I have 1 specimen borrowed from the U. S. N. M.) but the prothorax of the Haitian species is less inflated at sides below the margin, and the 2nd elytral stria is not abbreviated basally as it is in Isevistriatus. Moreover the latter lacks scutellar striae and is much more alutaceous across the base of the elytra. The striae of the elytra are deeper at apex in the Haitian species.
Tachys (Tachyta) noctis n. sp.
Form average for Tachyta, rather parallel and subde- pressed, but head smaller than usual; black, appendages, mouth parts, and lateral margins of prothorax and (less distinctly) of elytra testaceous; upper surface entirely moderately alutaceous. Head 2/3 width prothorax; eyes only slightly prominent; front with usual 2 weak impres- sions; antennae short, 2nd and 3rd joints subequal, outer ones (except apical) moniliform ; mentum not biperforate, toothed at middle. Prothorax 2/5 or slightly less wider than long; sides slightly rounded anteriorly, approximately straight before ,the posterior angles (sometimes minutely, faintly sinuate) ; posterior angles nearly right; fine but distinct costa each side from base for 1/3 of length within (not on) margin ; lateral margins translucent, evenly ex- planate from base to apex, as wide or wider than in T. flovicauda; middle line and basal transverse impression well marked, anterior impression very faint. Elytra only slightly depressed; each about &striate (7th and 8th striae



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19351 West Indian Curabidse 176
barely detectable), inner strise strongly impressed, outer shallower, all except sutural slightly abbreviated at extreme apex; apical stria recurved as usual: outer edge 3rd inter- val with setigerous punctures just before middle and %, from apex. Two basal joints 8 front tarsus dilated, with anterior apical angles somewhat produced. Length 2.25- 2.5 ; width 1.0 mm. (slightly more or less). HAITI: holotype (M. C. Z. no. 22014) and 4 paratypes from Roche Croix, Massif de la Hotte, 5,000 ft, Oct. 13; 3 paratypes from northeastern foothills La Hotte, Oct 10- 24 ; under bark.
Resembles in color the description of marginicollis Schaum (now called neotropicus Csihi) of Venezuela except that the appendages, of fwctis, are testaceous (not rufo- piceous), with femora not infuseate. The elytra are described as bistriate in wwrginicollis; there are numerous striae in noetic.
Tachys (Tachyura) tritax n. sp.
Form average for subgenus, convex; piceous, not dis- tinctly spotted, shining, moderately iridescent ; appendages and mouth parts testaceous. Head about % width pro- thorax, eyes prominent; front shining, briefly bi-impressed anteriorly; antennae moderate, middle joints about longer than wide; menturn not biperforate, toothed at middle. Prothorax about wider than long, sides strong- ly arcuate anteriorly, slightly sinuate before the obtuse but not blunted basal angles; latter briefly carinate; side margins narrow; basal transverse impression deep, punc- tulate, but not distinctly foveate; middle lime very fine, anterior impression nearly obsolete. Ely tra each 3-striate ; sutural stria entire and recurved as usual, 2nd abbreviated about 1/6 from base and apex, 3rd extending from anterior puncture (about 1/3 from base) to slightly behind posterior puncture (about 1/3 from apex), all 3 striae well impressed- Male with 2 baaal joints each front tarsus slightly dilated, with anterior apical angles slightly pro- duced. Length 2.2-2.7; width 1.0-1.1 mm. HAITI : holotype 8 (M. C. Z. no. 22015) and 5 paratypes from Camp Pen-in, Oct. 8-27; 10 paratypes from Trou Caiman, Nov. 15-20; 1 paratype from Emery, Sept. 6; 2



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176 Psyche [December
paratypes from Mt. Trou d'Eau, 4,000 ft., Nov. 19; most taken beside temporary muddy pools in roads. In Hayward's key to the North American species of Tachyura (Trans. American Ent. Soc. 26, 1899, pp. 202-), tritax runs to incurvus Say, agreeing in general form and in the structure of the basal transverse groove of the prothorax, but incurvus is spotted and has only a sutural stria on each elytron. Most of the known Central Ameri- can species of this group have the basal groove strongly tri- or uni-foveate at middle, and none has other characters as in tritax. The striation of the elytra in the latter is remarkable for its constancy.
Tachys (s. s.) carib n. sp.
Moderately broad and depressed; rather shining testa- ceous or brownish testaceous, head darker, iridescence faint. Head about 2/3 width prothorax, but with eyes prominent, forming nearly right angles with sides of head behind them; antennae with middle joints 1/^ or slightly.more longer than wide; front with usual 2 rather shallow sub- parallel impressions; mentum biperforate and toothed. Prothorax about 3/5 wider than long; sides more or less strongly sinuate just before the right (but finely blunted) posterior angles ; disk with usual impressions. Elytra rather broadly oval; humeri rounded but distinct; each elytron with about 3 inner striae more or less impressed except at apex, outer strise faint or absent; anterior dorsal puncture almost on 4th stria 1/3 from base, posterior within hooked tip of recurved stride. Male with 2 basal joints each front tarsus narrowly dilated. Length 1.8-2.2 ; width 0.8-0.9 mm.
HAITI, CUBA, JAMAICA, PUERTO RICO: holotype 8 (M. C. Z. no. 22017) and 95 paratypes from Ennery, Haiti, Sept. 6-11; additional paratypes from Haiti as follows: 7, northeastern foothills La Hotte, 2000-4000 ft., Oct. 10-24 ; 10, Camp Perrin, Oct. 8-27; 1, Riviere Froide, Oct. 3. Also
the following specimens, not types : Cuba : 23, soledad, near Cienfuegos, Aug. 2-12; Jamaica: 1, Rio Cobre, 5 mi. above Spanishtown, Aug. 9; 1, Blue Mts., about 4,500 ft., Aug. 13-20. All specimens taken by myself by washing out gravel bars, piles of stones, and trash by clear, running



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19351 West Indian Carabidse 177
brooks and rivers. Also Puerto Rico: 2, Serralles Finca, Ponce, June 5, 1934, R. G. Oakley (U. S. N. M.) . In my key (1. c.) this runs to Tachys abruptus Darl., but carib is much smaller, with sides of prothorax more sinuate basally and dilated joints of 8 front tarsi much narrower.
Tachys trechulus n. sp.
Convex, rather ventricose, but otherwise of normal Tachys (s. s.) form ; piceous or rufo-piceous, with seri- ceous lustre but scarcely distinct iridescence ; appendages brownish testaceous, antennae darker in basal V^.. Head about 7/10 width prothorax, narrow and with elongate mandibles; eyes only slightly prominent; frontal sulci narrow, subparallel ; antennae moderate in length but with unusually long and conspicuous setae, middle joints about jh longer than wide; mentum biperforate and toothed. Prothorax just over 1/3 wider than long; sides arcuate anteriorly, straight posteriorly, very faintly sinuate before the obtuse, finely blunted posterior angles; base somewhat oblique at sides; disk convex; basal transverse and median longitudinal grooves fine, anterior transverse impression obsolete. Elytra oval, very convex, with narrowed, rounded humeri; each with entire sutural stria recurved as usual and several additional striae faintly indicated on disk; anterior dorsal puncture on 4th interval about 1/3 from base, posterior within hooked tip of recurved stride. Inner
wings vestigial, strap-like, about 2/5 length of elytra. Two basal joints 8 front tarsus moderately dilated, with an- terior apical angles produced. Length 2.6; width 1.2 mm. JAMAICA: holotype s (M. C. Z. no. 22018) and 1 paratype from Blue Mt. Forest Reserve, main range Blue Mts., 5,000-7,000 ft., Aug. 17-19 ; under deeply buried stones in damp forest.
Unique, among the Tachys
(s. s.) with 2 joints each 8
front tarsus dilated, in the atrophied wings and (conse- quently) ventricose elytra.
Perileptus dentifer n. sp.
Elongate, depressed ; rufo-testaceous to brownish piceous, legs paler, antennae scarcely so; upper surface pubescent



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178 Psyche [December
as usual. Head barely narrower than prothorax by meas- urement, appearing at least as wide ; eyes very prominent ; temples very briefly subparallel behind eyes ; front shining, lightly punctate; antennae slender, middle joints 3y2-4 times long as wide; mentum toothed as usual. Prothorax cordate, 14 or less wider than long; least width before basal angles slightly less than 2/3 greatest width; sides very strongly sinuate before basal angles ; latter very acute, directed almost laterally; sides of base oblique or some- what emarginate near angles; lateral margins entire, nor- mal, more or less crenate posteriorly; disk finely, moder- ately closely (not densely) punctate ; impressions as usual. Elytra rather finely and closely punctate; sutural stria entire except at base, 2 or 3 other striae faintly indicated, 3rd 3-punctate as usual. Male with 2 basal joints each front tarsus moderately dilated. Length 2.7-3.3 ; width 0.9-1.1 mm.
HAITI and PUERTO RICO: holotype
8 (M. C. Z. no. 22019)
and 43 paratypes from Ennery, Haiti, Sept. 6-11. Also the following specimens, not types : Haiti : 2, Riviere Froide, Oct. 3; 11, Camp Perrin, Oct. 8-27; 5, northeastern foot- hills La Hotte, 2,000-4,000 ft., Oct. 10-24; Puerto Rico: 2, Serralles Finca, Ponce, June 5, 1934, R. G. Oakley (U. S. N. M.). All my specimens were taken in gravel beside swift, clear streams.
Similar to the previously know West Indian Perileptus (cf. Darlington, Psyche 41, 1934, pp. 86-88) and like them a member of the areolatus group, but easily known by the very acute posterior prothoracic angles. Perikptus minutus n. sp.
Slender, depressed ; testaceous or rufo-testaceous, rather shining ; pubescence as usual. Head barely narrower than prothorax; eyes prominent; temples very briefly sub- parallel behind eyes; front obsoletely punctate, shining; antennae relatively stout, middle joints 2-2y2 times long as wide ; mentum toothed. Prothora,~ subcordate, about 1/8 wider than long, sides narrowed, sinuate, and then subparallel before right posterior angles ; latter not truly basal, set slightly forward from the base so that prothorax is very briefly pedunculate; width across basal angles 2/3



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19351 West Indian Carabidee 179
or slightly more greatest width; width basal peduncle 1/^ greatest width ; side margins narrow, entire, slightly crenate basally; disk with usual impressions, finely, not densely punctate. Elytra, rather closely punctate, punc- tures relatively much coarser than in other American Perileptus, sutural stria entire except at base, other striae scarcely indicated; usual 3 setigerous punctures on posi- tion of 3rd stria about 1/5 from base, behind middle, and about 1/10 from apex. Male with 2 basal joints each front tarsus dilated as usual. Length 1.8-1.9; width about 0.5 mm.
JAMAICA and HAITI: holotype s
(M. C. Z. no. 22020) and
1 9 paratype from Rio Cobre, 5 mi. above Spanishtown, Jamaica, Aug. 29 ; 1
? paratype from Ennery, Haiti, Sept.
6; all washed from bars of river gravel. Also a member of the areolatus group of Perileptus, but very distinct within the group because of its small size, rather stout antennae, relatively coarsely punctate elytra, and subpedunculate prothorax.
Dyschromus tiburonicus n. sp.
Form average; dull black with a brownish, purplish cast especially on the elytra; mouth parts and appendages brownish to piceous. Head about 2/3 width prothorax; eyes moderate; front dull but not punctate, with 2 short impressions between anterior edges of eyes; vertex lightly transversely impressed ; antennae as usual in genus ; mentum with short, emarginate tooth; labial palpi both sexes with apical joint very broadly triangular, apex nearly as wide as inner side in $ ; apical joint maxillary palpi slender, nar- rowly truncate. Prothorax about 1/3 wider than long, slightly narrowed behind; sides straight or very broadly and slightly sinuate before right or slightly obtuse basal angles; base and apex not margined, side margins narrow, each with a seta 1/3 from apex and at basal angle; disk rather flat basally, more finely alutaceous than head, not punctate ; transverse impressions rather vague ; middle line fine but well impressed except at base and apex; basal fovese linear, about 1/3 length prothorax, scarcely nearer sides than middle. EZytra convex, opaque; striae entire, moderately impressed, impunctate except for a strong punc-



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180 Psyche [December
ture almost at base 2nd stria; first stria sometimes briefly obliterated near base, leaving 1 or 2 punctiform or linear impressions basally ; intervals slightly convex, not irregular nor interrupted. Abdomen with last 3 segments trans- versely grooved at base as usual, grooves subcrenate lat- erally; prosternal process strongly margined; mesepister- num with a cluster of punctures- near middle, lower surface otherwise impunctate. Male with anterior tarsi moderately dilated, 4 joints of each biseriately squammulose. Length 10.5-11.5 ; width 4.1-4.6 mm.
HAITI: holotype 8 (M. C. Z. no. 22021) and 3 s para- types from northeastern foothills La Hotte, 2,000-4,000 ft., Oct. 10-24; 1 $ paratype from north slope La Hotte itself, 5,000 ft., Oct. 18; all under stones and logs in damp woods and coffee plantations.
This species is very distinct from the two Dyschromus previously known from Haiti (opacus Chd. and cupripemis Chd.), both of which we possess, in having the elytra with impressed striae and regular intervals.
It is perhaps more
like the Mexican chrysophanus Bates, known to me only by description, but the latter has an unmargined prosternal process and probably has shallower elytral striae. Loxandrus mutans n. sp.
Rather slender and subparallel, not much depressed; black; antennae (especially basally), mouth parts and tarsi brownish testaceous ; lower surface and legs brownish or rufo-piceous; head and prothorax moderately shining, elytra dull, alutaceous in both sexes, but especially so in s . Head about % width prothorax; mentum tooth broad, sub- truncate apically, slightly impressed along middle. Pro- thorax about VA, wider than long; sides slightly to moder- ately arcuate anteriorly, moderately narrowed and sinuate posteriorly before right posterior angles; margins very narrow, evenly reflexed ; disk moderately convex ; middle line fine but distinct except at base and apex; transverse impressions subobsolete ; fine anterior submarginal line widely interrupted at middle; basal foveae linear, rather deep, at least 1/3 length prothorax ; surface impunctate ex- cept vaguely punctate at middle near base. Elytra narrow and not much depressed, alutaceous, with fine, entire, mod-



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19351 West Indian Carabidee 181
erately impressed, finely punctulate strise; inner intervals slightly convex, outer nearly flat; single dorsal puncture on inner side 3rd interval near middle. Metepisterna about twice as long as wide, narrowed posteriorly. Hind tarsi sulcate both sides above; 5th tarsal joint with about 3 setse each side below. Male anterior tarsi each with first 3 joints conspicuously dilated and oblique. Length 8-9 ; width 2.7-3.3 mm.
HAITI: holotype 8 (M. C. Z. no. 22022) and 19 paratypes from Etang Lachaux, Oct. 26-27; 10 paratypes from swamps north of Dessalines, Sept. 11; 1 paratype from Damien, near Port-au-Prince, A. Audant collector. My specimens were taken under trash and in loose soil a little above water level beside standing water. This species belongs to a small group of Loxandrus char- acterized by the elytra being opaque, especially in the ?. From opa-eulus Bates of Brazil, mutans differs in having the body piceous, not clear red, below; from seulptilis Bates of Mexico and Panama, in having the sides of the prothorax sinuate posteriorly. Both Bates' species are unfortunately known to me only by description.
In my first West Indian paper (pp. 92-93) I gave a key to the 15 insular species of Colpodes then known. Aquisi-
tion of more material has now more than doubled the num- ber of species, so that a new key is necessary. In compos-
ing it, I have avoided as much as possible using the form of .
the metepisterna as a character, for its use results in a very unnatural classification, full of borderline species, and very difficult to use. In deference to the past classifications of Chaudoir and Bates, however, I have arranged the couplets in such a way that all species the names of which occur in the key up to and including couplet 21 have the metepi- sterna with outer edge (not including the posterior lobe which overlaps the first ventral) not longer by measure- ment than 1% times the length of the anterior edge. These species would fall in Chaudoir's groups I or I1 in his 1878 revision (Ann. Soc. Ent. France (5) 8, pp. 278-382). Species named in couplet 22 or below have the metepisterna with outer edge not less than 1% times the anterior, and



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182 Psyche [December
would fall in Chaudoir's group 111. No additional material has been secured from the Lesser Antilles, where there are still 3 species recorded which are unknown to me, so the following key has been constructed to cover only the species of the Greater Antilles. In this region there is now only one described species which I do not know, C. (Metalloso- mus) cupraseens Mots. (Bull. Soc. Nat. Moscou 37, 1864, no. 4, p. 305), described from "St. Domingue aux Antilles." The description is superficial, but the species is evidently very different from any I have seen.
With the exception of the widely ranging C. asquinoctialis Chd., I know of no Colpodes which occurs either on the Greater Antilles and the mainland (I have seen at least half the known Central American species), the Greater and Lesser Antilles, any two of the greater islands, or even upon any two sepaxate mountain ranges on a single island. They are evidently so local that I feel safe in describing West Indian species without discussing their relationship individually with the continental fauna. Key to Greater An,tiZZea/n Colpodes (excepting cupraseens Mots.)
Tibiae finely sulcate along extreme outer edge ; tarsi strongly trisulcate above .................... .27. Tibise not sulcate on outer edge; tarsi with not more than 2 sulei above ........................... .2. One or more supra-ocular and/or lateral thoracic setas missing ; Jamaican species ................... .3. . Two supra-orbital and 2 lateral thoracic setae each side; species not on Jamaica, except for the widely dis- tributed asquinoctiaZ<s ........................ .8. Two supra-ocular setae each side ................. .4. One supra-ocular seta (the anterior) missing ..... .5. 4. Anterior lateral thoracic seta present, posterior miss- ing; elytra much duller than head and prothorax. .. cinchonas Darl.
Anterior seta missing, posterior present; elytra not duller than head and prothorax ; (metepisterna much more elongate) ............................ .22.



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19351 West Indian Carabidse 183
5. Anterior thoracic seta present, posterior missing ; ............ elytral striae subcrenulate faber n. sp.
Anterior seta absent, posterior present; elytral striae ......................................
smooth 6.
......... 6. Broad, prothorax about 1/3 wider than long vagepunctatus Darl.
Sender, prothorax as wide as to 1/10 wider than long 7. 7. Elytra normal ; prothorax more subquadrate, posterior ...... angles obtuse, narrowly rounded
macer Darl.
Elytra oval, subventricose ; prothorax oval with very broadly rounded posterior angles .... subovalis n. sp. 8. Length 6-8 mm. ; form very Agonum-like ......... .9. ...................................
Size larger 10.
9. Sides of prothorax not sinuate; elytral striae relatively shallower ........................ agonellus n. sp. Sides of prothorax sinuate before base; striae deeper. . pavens n. sp.
10. Posterior marginal seta on each side of prothorax about 1/10 prothoracic length before basal angles ; elytra duller than head and prothorax ......... .11. Said seta on or not more than 1/20 before basal angles ; elytra not duller than head and prothorax ..... .13. 11. Eyes small, scarcely more prominent than gense; pro- thorax not margined at middle anteriorly. ........ marcus n. sp.
Eyes larger, much more prominent than gense; pro- thorax with fine, entire anterior margin ....... .12. 12. Elytral striae moderately fine; s hind trochanters 1/3 length femora, unmodified .......... .amone n. sp. Striae extremely fine and superficial ; 8 hind trochant- ers 2/5-l/9, length femora, bluntly pointed ......... wolla n. sp.
13. Hind tarsi with 4th joint emarginate, outer lobe formed about like inner and not much longer; habits not riparian .................................... 14. Hind tarsi with 4th joint emarginate, outer lobe much longer than inner; species associated with running water ...................................... 26. 14. Elytral striae interrupted ...................... .23. Elytral striae (except rarely the outer ones) not inter- rupted ...................... ............... 15.



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184 Psyche [December
15. Prothorax not narrowed basally, base broad as middle ..................................... 24. Prothorax more or less narrowed basally ........ .16. 16. Metepisterna with outer edge not over v2 longer than anterior .................................... 17. Outer edge metepisterna about 2 )( length anterior .......................................
edge 25.
17. Head relatively small, less than 2/3 width prothorax, genae not prominent, oblique, barely convex in pro- file; mentum tooth finely pointed (northern Haiti) 18. Head larger, about 7/10 or more width prothorax, gense convex and prominent; mentum tooth blunt or .................
emarginate (southern Haiti) .19.
18. Prothorax subquadrate with posterior angles almost right; not margined at middle anteriorly; s poste- rior trochanters acuminate, 3/5 length femora ..... christophe n. sp.
Prothorax with sides much rounded and posterior angles obtuse ; anterior margin fine but entire ; s pos- terior trochanters probably not acuminate ......... cychrinus n. sp.
19. More slender and depressed ; prothorax finely margined anteriorly (see also under description) (La Hotte) . . constricticeps n. sp.
Stouter ; prothorax not margined at middle anteriorly .................................
(La Selle) .20.
20. Larger and broader, with head relatively narrower; margins of prothorax moderately to broadly ex- planate .................................... 21. Slightly smaller and narrower, head relatively broader; margins of prothorax rather narrowly explanate (see description for final identification)
.... tipoto n. sp.
21. Prothorax subquadrate, base VA, or more wider than apex, margins moderately explanate ; posterior dorsal puncture of elytra present ............ jsegeri Dej. Prothorax subcordate, base 1/10 to 1/7 wider than apex, sides rather widely explanate ; posterior elytral puncture missing ................ subcordens n. sp. 22. Brown ; external elytral striae punctulate . punctus n. sp. Bluish purple; striae not punctulate. ..... bruesi n. sp. 23. (No alternative) .................. fractilinea Darl.



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19361 West /ndio,~ Carabidss + 186
24. (No alternative) ..................... .visitor n. sp. 25. (No alternative) .................... baragm n. sp. 26. Elytra bluish or greenish, striae not interrupted. ..... ssywinoctwiis Chd.
Elytra not metallic, striae interrupted. dtift~tminis n. sp- 27. (No alternative) ................... .latelytra n. sp. Elongate, moderately convex ; brownish piceous, lower surface and appendages rufescent ; not strongly shining, but elytra not much duller than head and prothorax. Head large, 4/5 or slightly more width prothorax; eyes not prominent, genas only slightly converging posteriorly; neck very wide, slightly narrower than gense; ante- rior supra-ocular seta missing; antennae short, not reach- ing much beyond base prothorax; menturn tooth blunt at apex. Prothorax subcordate, about 1/4, wider than long; base squarely truncate, about equal to or a trifle narrower than apex; sides broadly, not strongly arcuate in anterior %, moderately strong-Iy sinuate before the right but finely blunted posterior angles, margins rather broadly explanate for such a slender species, posterior marginal seta missing; base and apex finely margined, latter sometimes rather in- distinctly so; disk with usual impressions; basal fovese deep, not sharply defined,
each with an impressed area
extending forward parallel to margin almost to apex, not punctate.
Elytra slightly (^,-3/10) wider than prothorax, rather elongate, widest about 1/3 from apex; humeri dis- tinct; sides slightly converging anteriorly, faintly sinuate 1/5 or 1/6 from base and before apices, which are rather broadly, subindependently rounded; disk moderately con- vex ; striaa well impressed, rather coarsely punctate or sub- crenate, with individual punctures not well defined ; inter- vals convex, 3rd either without visible dorsal punctures or with single one at or behind middle. Metepisterna with outer edge slightly longer than anterior, not or only faintly margined at sides; body impunctate below. Tibise not grooved on outer edge; hind tarsi not or indistinctly grooved above, 4th joint emarginate, outer lobe very slightly longer than inner; front tarsi not grooved above. Length 11.5- 13 ; width 3.7-3.8 mm,




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186 Psyche [December
JAMAICA: holotype (M. C. Z. no. 21992) and 3 paratypes from Blue Mt. Forest Reserve (main range Blue Mts.), 5,000-6,000 ft., Aug. 17-19, 1934; under stones and chips in wet forest. Unfortunately all are
s s . The species is very
isolated, taxonomically.
Colpodes subovalis n. sp.
Slender anteriorly, but rather ventricose ; piceous, lower surface and legs a little reddish, antennae and palpi testa- ceous, former infuscate on 3 basal joints ; rather shining, elytra not duller. Head elongate and slender, less than %, width prothorax; eyes moderately prominent, genae oblique, not prominent ; anterior supra-ocular seta missing ; antennae slender, 7th joint reaching about to base pro- thorax; mentum tooth broad, blunt or almost pointed. Prothorax rather small, 1/10 or less wider than long, sub- oval, slightly variable in outline, broadest at or behind middle; sides arcuate from base to apex or nearly straight and converging anteriorly and (less distinctly) posteriorly ; basal angles broadly rounded, anterior angles better de- fined but scarcely prominent; margins rather narrowly ex- planate; posterior marginal seta a little before angle, an- terior missing; disk; convex, impressed as usual, basal foveae moderately deep but not sharply defined, each with a vague depressed area extending forward nearly to apex; base and apex finely margined; surface slightly roughened basally but not distinctly punctate. Elytra much (about %,) wider than prothorax, suboval, widest behind middle ; sides straight and converging anteriorly; humeri broadly rounded; margins slightly sinuate before apices, which are independently pointed but not much produced ; striae deep, not punctate; intervals convex, 3rd 3-punctate as usual. Metepisterna with outer margins fully & longer than an- terior; mesepisterna somewhat punctate, lower surface otherwise impunctate. Tibiae not sulcate on outer edge; hind tarsi grooved each side above, less strongly internally, sulci not approximate, 4th joint emarginate, outer lobe much produced ; front tarsi not distinctly grooved. Length 12.5-14; width 4.2-4.7 mm.
JAMAICA: holotype 8 (M. C. Z. no. 21993) and 8 para- types from Portland Gap, Blue Mt. Forest Reserve, main



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19351 West Indian Caxabidse 187
range Blue Mts., about 5,000 ft., Aug. 17-19, 1934; in piles of dead vegetation and cracks in rotten logs. Related to C. macer Darl. of Cinchona, Blue Mts., but differing notably in form.
Colpodes agonellus n. sp.
Small, broad, moderately depressed, very like an Agonum (s. str.) of the melanarium group; piceous to black, suture and margins of elytra sometimes reddish, appendages brownish; surface moderately shining, with microscopic silky alutaceous sculpture, elytra not duller. Head normal, between 3/5 and 2/3 width prothorax; eyes moderately prominent, genae short and oblique; both supra-ocular setae present; antennae moderate, apex 7th joint reaching about to base prothorax; mentum tooth triangular, narrowly rounded at apex. Prothorax rather broad, about '%. wider than long; base almost truncate, VA, (more or less) wider than apex; sides nearly evenly rounded except sometimes faintly sinuate before the obtuse but fairly distinct poste- rior angles ; margins translucent, narrowly explanate an- teriorly, broader posteriorly, each with usual 2 setae; base and apex finely margined; disk moderately convex, de- pressed at sides basally but without distinct fovese; middle line well impressed, transverse impressions vague ; surface not distinctly punctate. Elytra broad, not quite $$ wider than prothorax; humeri somewhat rounded but distinct; margins moderately sinuate before apices, latter subinde- pendently rounded or subtruncate, not produced; striae moderately impressed, impunctate ; 3rd interval 3-punctate, first puncture nearly on 3rd, second and third punctures on 2nd stria. Metepisterna with outer edges about longer than anterior ; inner wings dimorphic, full or reduced ; body below impunctate. Tibiae not sulcate on outer edge ; pos- terior tarsi sulcate each side above, sulci not approximate, 4th joint rather small, simply emarginate; anterior tarsi lightly or not distinctly sulcate. Length 6.5-7.5 ; width 2.7-3.3 mm.
HAITI: holotype 8 (M. C. Z. no. 21994) and 80 para- types from La Visite and vicinity, 5,000 (on Mt. Bourette) -7,000 ft., La Selle Range, Sept. 16-23, 1934; under various cover on the ground in cloud forest and damp gullies.




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188 Psyche [December
For comparisons, see under the following species. Colpodes pavens n. sp.
so similar to the preceding, C. agonellus, as to answer to the description of that species with the following changes : Color somewhat more brownish piceous. Head about 2/3 width prothorax. Prothorax relatively narrower, 1/3 wider than long; base 1/5 wider than apex; sides sinuate before the posterior angles, which are relatively prominent and approximately right, although more or less finely blunted ; margins finer. Elytra relatively a little narrower, 2/5 wider than prothorax; striae distinctly deep- er; intervals more convex. Length 6.5 (slightly more or less) ; width about 2.6 mm.
HAITI: holotype 8 (M. C. Z. no. 21995) and 3 paratypes, taken with the preceding species.
These species, agonellus and pavens, are among the most slightly specialized Colpodes, with the 4th joint of the posterior tarsus scarcely more emarginate, although some- what shorter and broader, than in, for example, Agonum melancwium Dej. of the United States. They recall the Mexican Colpodes of the nugax group, but the latter re- semble Anchomenus more closely than Agonum s. str. Colpodes marcus n. sp.
Rather elongate and convex, appendages shorter than average ; piceous, appendages scarcely paler ; head and prothorax moderately shining, elytra somewhat duller. Head large, 4/5 or slightly less width prothorax, but ap- pearing wider; eyes small, hardly at all prominent; genae oblique, only slightly convex in profile, neck very little nar- rowed ; 2 supra-ocular setae each side ; antennae rather short, reaching about to base prothorax ; mentum tooth triangular, pointed or somewhat blunted. Prothorax as long to about 1/8 longer than wide, subquadrate, slightly narrowed an- teriorly and posteriorly, base about wide as apex; sides slightly, broadly arcuate, faintly sinuate at posterior mar- ginal bristle; margins very narrow, each with usual 2 setae, the posterior 1/10 or more before base; posterior angles rounded, but not very broadly so; base finely margined, apex not; disk convex except depressed near posterior



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19351 West India-n Carabidse 189
angles to form shallow, poorly defined foveae, vaguely ex- tended anteriorly ; middle line distinct, transverse impres- sions weak; surface impunctate. Elytra rather narrow, about 1/3 wider than prothorax; humeri
distinct; sides
arcuate-parallel at middle, weakly sinuate before more or less independently rounded but not produced apices; striae rather lightly impressed, impunctate ; intervals barely con- vex, 3rd 3-punctate, punctures nearly on 3rd (1) and 2nd
(2 & 3) striae. Metepisterna with outer and anterior mar- gins about equal; lower surface impunctate. Tibiae not sulcate on outer edge; hind tarsi sulcate above externally, at most faintly so internally, 4th joint about rectangularly emarginate, not much lobed externally; front tarsi not dis- tinctly sulcate; 8 posterior trochanters more or less pointed, about v2 length femora. Length 10-13; width 3.7-4.0 mm.
HAITI: holotype 8 (M. C. Z. no. 21996) and 47 para- types from Mt. La Hotte, 5,000-7,800 ft., Oct. 16-17, 1934; 1 paratype from Roche Croix, on a spur of La Hotte, 5,000 ft., Oct. 13; under cover on the ground and in rotten logs in wet cloud forest.
I have named this species for my cook and number one boy on my La Hotte trip, Marc Jacques; the two following are named for my porters.
Colpodes wolla n. sp.
Rather elongate, convex, more lightly built than marcus ; piceous, antennae somewhat paler, legs scarcely so; head and prothorax moderately shining, elytra rather stongly alutaceous. Head 2/3 or slightly more width prothorax; eyes rather small but somewhat prominent; genae sub- oblique, slightly convex but not very prominent; 2 supra- ocular set= each side; antennae average, 9th joint reaching about to base prothorax; mentum tooth pointed or slightly blunted. Prothorax subquadrate, as long or slightly longer than wide, slightly narrowed posteriorly and anteriorly; base slightly wider than apex; sides broadly and evenly rounded, sometimes faintly sinuate posteriorly; margins narrowly explanate, each with usual 2 setse, the posterior about 1/10 before base; posterior angles obtuse, narrowly rounded ; base finely margined, apex variably so ; disk very



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190 Psyche [December
convex, almost gibbous except broadly depressed near pos- terior angles to form rather poorly defined but deep fovea?, which are continued forward parallel to margins as usual, not punctate; discal impressed lines as usual. Elytra a little (about 3/10) wider than prothorax, a little less parallel than in marcus, very convex; humeri distinct; subapical sinuation slight; apices more or less independent- ly narrowly rounded; striae excessively fine but entire; in- tervals flat, 3rd with 3 dorsal punctures as in preceding species. Metepisterna with outer and anterior edges about equal; lower surface impunctate. Tibiae not sulcate on outer edge; posterior tarsi above lightly grooved externally, not or faintly internally, 4th joint moderately deeply emarginate, outer lobe a little longer than inner; front tarsi not distinctly sulcate; 8 posterior trochanters some- what pointed, about % length femora. Length 11-14; width 3.8-4.4 mm.
HAITI, MASSIF DE LA HOTTE: holotype
$ (M. C. Z. no.
21997) and 3 paratypes from ridge just above Roche Croix, on a spur of Mt. La Hotte, 5,000 ft., Oct. 13; 1 $ from N. slope La Hotte, 5,000 ft., Oct. 14; 1 Q from Des- barrikre, about 4,000 ft., Oct. 13; 2
8 Q from first rain
forest on auto road N. of Camp Perrin, about 3,000 ft., Oct. 21; all taken in loose leaf mold and under loose bark of dead trees in wet forest.
Colpodes amone n. sp.
So similar to the preceding, wolla, as to answer to the description of that species except as follows: Smaller and more graceful; slightly less convex. Head relatively a trifle wider; antennae with 7th joint reaching about to base prothorax. Prothorax with fine anterior margin more dis- tinct. Elytra scarcely or not sinuate on outer margin be- fore apex (sometimes scarcely so in wolla) ; striae slightly more distinct but still fine. Posterior trochanters of 8 short, rounded-truncate as in Q . Length 10-11.5 ; width 3.7-3.9 mm.
HAITI: holotype 8 (M. C. Z. no. 21998) and 7 paratypes from on and near summit La Hotte, about 7,000-7,800 ft., Oct. 16-17, 1934; in moss and loose leaf mold in forest.



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19351 West Indian Carabidss 191
Colpodes christophe n. sp.
Very large, rather broad and depressed ; black, moderate- ly shining, elytra very slightly duller; appendages piceous. Head barely more than 3/5 width prothorax; eyes small but somewhat prominent; gene oblique, slightly convex, not prominent; 2 supra-ocular sebe each side; antennae rather short, 9th or 10th joint reaching base prothorax; menturn tooth finely pointed. Prothorax large, subquad- rate, 1/5 or 1/6 wider than long; base squarely truncate, not quite wider than apex; sides slightly arcuate ante- riorly, broadly, slightly sinuate before posterior angles, which are almost right, with apices very narrowly rounded ; side margins broadly but not strongly explanate, each with 2 setae; base finely margined, apex not; disk rather flat, irnpunctate, depressed posterio-laterally to form moderately deep, not very well defined fovese which extend anteriorly to or before middle; other impressed lines as usual. EIytra not much ( s not quite 1/5, a not quite 3/10) wider than prothorax, moderately convex ; sides arcuate-subparallel at middle, slightly sinuate before apices, which are nar- rowly, subindependently rounded; humeri distinct; striae somewhat impressed, entire, not punctate except for usual 3 dorsal punctures attached to 3rd interval ; intervals some- what convex. Metepisterna with outer edge somewhat longer than anterior; lower surface not punctate. Tibise not sulcate on outer edge; posterior tarsi finely grooved above externally, less strongly so internally, 4th joint mod- erately deeply emarginate, not distinctly lobed ; anterior tarsi finely grooved at sides in
s , not in 8 ; s posterior
trochanters about 3/5 length femora, suddenly narrowed just before middle of femoral length, thence very slender to apex ; $ trochanters slightly less than $$ length femora, but with apices subtuberculate at point corresponding to 8 apical process. Length 17-18; width 6-6.2 mm. HAITI: holotype 3 (M. C. Z. no. 22001) and 1 9 para- type from summit plateau of Mt. Basil, northern Haiti, 4,700 ft., Sept. 9, 1934; under stones in a tiny garden re- cently cleared from cloud forest.
This is one of the largest and finest species of the whole great genus Colpodes.




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192 Psyche [December
Colpodes cychrinus n. sp.
Rather large, broad, subdepressed ; black, moderately shining, elytra scarcely duller, appendages red-piceous. Head relatively narrow, between and 3/5 width pro- thorax, structure as in christophe. Prothorax wide, about 45/100 wider than long; base truncate, about 1/3 wider than apex; sides rather strongly arcuate throughout, with margins even broader and more strongly explanate than in christophe; posterior angles obtuse, but only very narrowly rounded; base and apex finely margined; disk about as in christophe. Elytra, not much (14, more or less) wider than prothorax, rather short, not or faintly sinuate near apex; striae a little shallower, intervals flat or nearly so; otherwise as in christophe. Characters of lower surface and legs ( Q ) as in christophe except hind trochanters ( $ ) only about 1/3 length femora, broad and not tuber- culate at apex.
HAITI: holotype Q (M. C. Z. no. 22002) and 2 $ para- types, all with the same data as the types of christophe. Closely related to christophe, but the prothorax is strik- ingly different in shape, and the hind trochanters are not modified in the Q and probably not in the 8, for in the three species I know with modified 8 trochanters (chris- tophe, marcus, and wolla) there is a corresponding but slighter modification in the 9.
Colpodes jaegeri (Dej .)
Dejean 1831, Spec. Coleop. 5, 728 (Anchomenus) Chaudoir 1859, Ann. Soc. Ent. France (3) 7,315 Chaudoir 1878, 1. c. (5) 8,280 & 300
Darlington 1934, Psyche 41, 92 (in key)
HAITI: summit of Morne Tranchant, Massif de la Selle (but not main range), about 6,000 ft., Nov. 11, 1934; 4
specimens, both sexes.
These agree very well with Dejean's
and Chaudoir's (1859) long descriptions. Colpodes subcordens n. sp.
Moderately broad, average convexity ; black, moderately shining, elytra not duller (whole surface sometimes ob- scured by a thin,
tenaceous coating of foreign matter) ;




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1~351 West Indim Carabidas 193
appendages red-piceous. Head about 2/3 width prothorax, formed as in jsgeri, with gem very convex in profile, scarcely less prominent than eyes; 2 supra-ocular setae each side; antennae average, apex 8th joint reaching to about base prothorax; menturn tooth variable, with apex bluntly rounded, aubtruncate, or marginate, channeled along middle. Prothorax subcordate, 1/3 or slightly less wider than long, base only slightly (1/10-1/7) wider than apex; sides strongly, often rather irregularly rounded anteriorly, very slightly to moderately sinuate before the slightly obtuse, very narrowly rounded of blunted posterior angles; margins rather widely explanate, each with usual 2 setse; base finely margined, apex not distinctly so; disk with usual impressions, as in jasgen, impunctate. EMra as in jssgeri except a little more narrowed basally, with sides less parallel; humeri distinct; stri~ deep, impunctate; in- tervals convex, 3rd 2-punctate, about v4 from base and at middle, 3rd (posterior) puncture uniformly missing. Metepisterna with outer edges only slightly longer than anterior; lower surface impunctate. Tibiae not sulcate ex- ternally; posterior tarsi sulcate each side above, but inner sulcus sometimes indistinct, sulci not approximate, 4th joint moderately deeply emarginate but only slightly lobed externally; anterior tarsi not distinctly grooved at sides in either sex. Length 12-15 ; width 4.6-5.5 mm. HAITI: holotype 8 (34. C, 2. no. 22003) and 17 para- types from La Visits and vicinity, main La Selle range, 5,000-7,000 ft., Sept. 16-23, 1934 ; under stones and in damp moss and loose leaf mold in cloud forest. Colpodes tipoto n. sp.
Form about average, less broad than in jsegeri and sub- cordons and more subparallel; piceous black, lower surface and appendages rufescent; moderately shining, elytra not duller. Head formed as described for swbcordens but rela- tively broader, about % width prothorax ; antennae slightly shorter ; menturn tooth blunted triangular. Prothorax sub- quadrate, % wider than long; base truncate, about 1/10 wider than apex; aides less rounded than in preceding species, straight or faintly sinuate before obtuse but dig- tinct basal angles; margins narrowly explanate, each with



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194 Psyche [December
2 setae; base finely margined, apex not; disk about as in jsegeri and subcordens. Elytra slightly (barely over 14) wider than prothorax; sides subparallel, faintly narrowed in basal 1/^ ; humeri distinct; sides slightly sinuate before apices ; latter narrowly subindependently rounded ; striae moderately deep but less so than in 2 preceding species, im- punctate ; intervals moderately convex, 3rd with anterior and middle punctures present, posterior missing. Mete- pisterna and lower surface as in subcordens, as are also tibia1 and tarsal characters. Length 10.5-11; width be- tween 3.5 & 4.0 mm.
HAITI: holotype 8 (M. C. Z. no. 22004) and 1 ? para- type from vicinity La Visite, main range La Selle, about 6,500 ft., Sept. 16-23, 1934; under deeply buried logs in cloud forest.
Named for the headman of the porters whom ton Bates, Mr. Andre Audant, and I had with La Selle trip.
Dr. Mars-
us on our
Colpodes constricticeps n. sp.
Rather slender, subdepressed ; piceous, lower surface and appendages a little more rufescent; moderately shining, elytra not duller. Head large, about 4/5 width prothorax; eyes small but slightly prominent; gens convex, prominent, head behind them rather strongly constricted (for Col- podes) at sides and across occiput; 2 supra-ocular set= each side; antennae moderate, 8th joint about opposite base prothorax; mentum tooth rather broad, apex slightly, rather broadly emarginate (probably variable) . Prothorax elongate-cordate, just over 1/10 wider than long; base squarely truncate, as wide as apex; sides arcuate anterior- ly, broadly sinuate before right posterior angles; margins narrowly explanate, each with usual 2 setae; base and apex finely margined; disk as in js&ge~i or subcordens except transverse impressions rather better defined and surface at extreme base (behind posterior transverse impression) irregularly transversely subrugose. Elytra elongate, about 2/7 wider than prothorax, somewhat depressed (much more than in jsegeri and allies), widest about 1/3 from apex; sides almost straight and converging in more than anterior 1/9; humeri distinct but narrow and rather rounded,



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1935] West Indian Carahidse 195
humeral marginal line strongly annulate (arcuate in jsegeri and allies) ; subapical sinuation slight; apices inde- pendently narrowly rounded ; striae moderately impressed, impunctate; intervals a little convex, 3rd with 2 punctures, near middle and VA, from apex, basal puncture missing. Lower surface and characters of tibiae and tarsi as described for subcordens, except 4th joint hind tarsi not at all more lobed externally than internally. Length 11 ; width 3.6 mm. HAITI: holotype 8 (M. C. Z. no. 22005), unique, from summit Mt. La Hotte, about 7,800 ft., Oct. 16, 1934; under a deeply buried stone in damp, mixed forest. A very distinct species, though related to the jsegeri group. The absence of the anterior dorsal puncture of elytra is probably not a character of much significance, for the puncture is absent in one specimen of my series of subcordens and is known to be either present or absent in at least one other Colpodes, haptoderoides Bates of Mexico (Trans. Ent. Soc. London 1891, p. 252).
Colpoldes punctus n. sp.
Moderately elongate and depressed ; dark brown, append- ages a little paler; rather shining, elytra not duller. Head 415 width prothorax; eyes moderately prominent; gense oblique, not prominent; 2 supra-ocular setse each side; an- tennse moderately elongate, 7th joint about opposite base prothorax; inentum tooth slender, pointed. Prothorax rather small, rounded-subquadrate, barely wider than long ; base very slightly lobed at middle, rounded-oblique at sides, slightly wider than apex; sides evenly, not strongly rounded, margins very narrowly explanate, anterior lateral seta missing ; basal angles rounded-obtuse ; base and apex finely margined; disk with middle line fine but distinct, transverse impressions vague ; basal f ovese moderate, rounded, with usual vague anterior extension, punctate-rugose, base of pronotum behind transverse impression also rugulose. Elytra, much (about 3/4) wider than prothorax, moderately elongate ; humeri distinct ; sides nearly parallel at middle, slightly sinuate before apices, which are irregularly, almost conjointly rounded ; disk rather depressed ; striae moderate, inner ones faintly, outer ones much more distinctly punctu- late; intervals flat or slightly convex, 3rd 3-punctate.



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196 Psyche [December
Metepisterna very elongate; they, sides of metasterna, mesepisterna, first ventral, and a small part of sides of prosternum closely, not very coarsely punctate. Tibiae not sulcate on outer edge; hind tarsi not distinctly grooved above, 4th joint deeply emarginate, strongly lobed exter- nally; front tarsi exceptionally broad in both sexes, not distinctly sulcate above. Length 11.5-13 ; width 4.2-4.6 mm. JAMAICA: holotype $ (M. C. Z. no. 22006) and 6 para- types from Cinchona (Blue Mts.), 5,000 ft., Jan. 1912, C. T. Brues collector.
Colpodes bruesi n. sp.
Elongate ; purplish blue-black, metallic color stronger on elytra, lower surface and legs rufopiceous, antennae dark; surface rather shining, elytra not duller. Head about % width prothorax, elongate ; eyes prominent ; gense oblique, not prominent; 2 supra-ocular setae each side; antennae slender, 6th or 7th joint about opposite base prothorax; menturn tooth acutely pointed. Prothorax barely wider than long, subquadrate oval; base about 1/3 wider than apex, formed as in punctus; sides evenly, not strongly rounded, margins moderate, much wider than in punctus, anterior seta missing ; basal angles obtuse, almost rounded ; base and apex finely margined; disk almost as in punctus except basal foveae more distinctly extended forward ; foveae and base rather lightly punctate. Elytra much (3h) wider than prothorax, disk rather depressed ; humeri dis- tinct; sides slightly convergent in anterior s, slightly sinuate before the apices, which are independently sub- angulate ; strise rather fine, impunctate ; intervals faintly convex, 3rd 3-punctate. Metepisterna very elongate ; sides of body below rather indistinctly punctate. Tibia1 and tarsal characters as in punctus. Length 15.5; width 5.5 mm.
JAMAICA: holotype 8 (M. C. Z. no. 22007), unique, from Newton, 3,000 ft., Jan., 1912, C. T. Brues collector. Colpodes visitor n. sp.
Broadly subfusiform; rather shining black or piceous, elytra sometimes faintly aeneous, lower surf ace and legs rufopiceous, mouth parts and antennae rufous. Head small,



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19351 West Indian Carabidz 197
a trifle less than 2/3 width prothorax; eyes prominent; genz short, oblique or slightly convex in profile; 2 supra- ocular set= each side; antenn~ average, 8th joint reaching opposite or beyond base prothorax; mentum tooth a little bluntly pointed. Prothorax about 113 wider than long, not narrowed behind, strongly so in front; base squarely trun- cate, more than y2 wider than apex; sides parallel in basal x, arcuate-converging anteriorly ; margins narrowly ex- planate, each with 2 seb; basal angles right, scarcely blunted; base and apex finely margined ; disk with middle line fine but distinct, transverse impressions vague; basal f ove~ rather deep, irregular, impunctate, vaguely prolonged anteriorly as usual. Elytra moderately (451100) wider than prothorax, rather short ; humeri distinct ; sides parallel at middle, sinuate before the independently rounded apices ; striz moderately deep, inner ones impunctate, outer irregu- lar, faintly punetulate or interrupted; intervals somewhat convex, 3rd 3-punetate as usual. Metepisterna with outer edges fully y2 longer than anterior; body below not punctate. Tibi~ not grooved on outer edge; hind tarsi sul- cate each side above, sulci not approximate, 4th joint mod- erately emarginate, scarcely lobed externally ; front tarsi not distinctly grooved above. Length 9-10.5; width 3.7- 4.1 mm.
HAITI: holotype 8 (M. C. Z. no. 22008) and 13 para- types from La Visite and vicinity, La Selle Range, about 6,000 ft., Sept. 16-23, 1934; in cracks in rotten logs and under loose bark and similar cover in cloud forest. colpodes baragua n. sp.
CoZpodes sp. Darl. 1934, Psyche 41, pp. 93 (in key) & 97 Form Anchomenus-like ; black, shining, appendages brownish piceous. Head about y4 width prothorax; eyes prominent; gem short and oblique ; 2 supra-ocular set= each side; antennze rather slender, 7th joint reaching be- yond base prothorax; mentum tooth with apex narrowly rounded. Prothomx cordate, wider than long, rather strongly narrowed in front and behind; base squarely trun- cate at middle, slightly oblique at sides, about 15/100 wider than apex ; sides strongly arcuate anteriorly, strongly sinuate before the right posterior angles ; margins rather



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198 Psyche [December
narrowly explanate, each with 2 set=; base and apex finely margined ; disk impressed as usual, basal fove~ relatively small and deep? impunctate. Elytra much (about 2/31 wid- er than prothorax; humeri distinct; sides subparallel, though slightly converging anteriorly in anterior $$, sinuate before the subindependently narrowly rounded apices; striz moderately deep, impunctate on disk, lightly punctate externally; intervals more or less convex, 3rd 3-punctate. Metepisterna elongate ; mesepisterna some- what punctate, lower surface otherwise impunctate. Tibi~ not grooved on outer edge; posterior tarsi sulcate each side above, sulci not approximate, 4th joint moderately emar- ginate, not more lobed externally than internally ; front tarsi vaguely sulcate.
Length 10.5; width about 4 mm.
CUBA: holotype 9 (M. C. Z. no. 22009), uniquz, from Baraguh (Camaguey), May 7, 1928, at light, L. C. Scara- muzza collector; from the collection of the Tropical Plant Research Foundation, now at the Harvard research station at Soledad. The specimen previously recorded by me was from Somorrostro (Havana), Barro collector. This species is strongly winged and found at low alti- tudes, and I at first thought it might occur also in Central America and have been previously described, but I cannot identify it with any Central American species. Colpodes =quinwtialis (Chd.)
Stenomemis jageri Mann.
Colpodes mannerheimi Chd.
A series which I took in Haiti and Jamaica is not dis- tinguishable from our specimens from Mexico, Panama, and Peru.
Colpodes altiflum,inis n. sp.
Moderately elongate; shining piceo~s~ lower surface and legs very slightly, antennze and palpi more distinctly rufescent. Head y4 width prothorax, formed as in zquinoctialis, with prominent eyes and oblique, not promi- nent genze ; 2 supra-ocular set2 each side ; antenn~ mod- erately slender? 7th joint reaching about to base prothorax; mentum tooth subtruncate or slightly emarginate at apex. Prothorax subcordate, % (more or less) wider than long,



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19351 West Ind,ian Cara bid= 199
formed somewhat as in zqui~octialis but with anterior angles much less prominent; base truncate at middle? rounded-oblique at sides? 1/10 or slightly more wider than apex; sides moderately, somewhat irregularly arcuate an- teriorly, briefly sinuate before posterior angles, which are right or slightly obtuse? very narrowly blunted ; explanate margins rather narrow, each with 2 set=; base finely mar- gined? apex rather indistinctly so; disk convex, with usual fine middle line and poorly defined transverse impressions ; basal foveze rather deep? round, only vaguely prolonged anteriorly? impunctate. Elytra considerably (not quite 2/3) wider than prothorax? of moderate length? shorter and with base more narrowed than in zquinoctialis, and with humeri more broadly rounded, though not entirely oblit- erated; sides moderately sinuate before apices? which are rather narrowly independently rounded? sometimes vaguely subangulate; stri~ fine? shallow, finely irregular and in- terrupted ; intervals flat or slightly convex, 3rd 3-punctate. Metepisterna with outer edges a little more than y2 longer than anterior edges? distinctly shorter than in aq~inoctialis~ although both species are winged; lower surface im- punctate. Tibiz not sulcate on outer edge; hind tarsi sul- cate each side above? sulci not closely approximate although tarsi are slender? 4th joint rather deeply emarginate? outer lobe much longer than inner; front tarsi more or less indistinctly sulcate above (these characters as in aquinoc- tidis, but all sulci less impressed in altijluminis). Length 11-11.5 ; width 3.8-4.0 mm.
HAITI: holotype 8 (M. C. Z. no. 22010) and 8 paratypes from vicinity of La Visite, main La Selle Range, about 6?000 ft.? Sept. 16-23? 1934; taken among loose stones and in piles of drift beside the stream called by the local Negroes the "Riviiire Blanche."
Colpodes latelytra n. sp.
Form about as in C. chalybeus Dej.? but elytra relatively wider; dark bluish black with faint greenish iridescence in some lights? moderately shining ; appendages black, Head rather small but with prominent eyes (not quite so promi- nent as in chalybeus), 516 width prothorax; genz short and oblique ; 2 supra-ocular setze each side ; antennze aver-



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ZOO Psyche [December
age, 8th joint about opposite base prothorax; mentum tooth narrowly blunted at apex. Prothorax small, subcordate but not strongly narrowed basally, about 118 wider than long; base truncate at middle, rounded-oblique at sides, about 115 wider than apex; sides arcuate in anterior v4, moderately sinuate before the right, narrowly blunted basal angles ; margins rather narrowly explanate anteriorly, somewhat broader posteriorly, each with posterior seta present, anterior missing (as in chalybeus) ; disk some- what convex, middle line and anterior transverse impres- sion fine but distinct, posterior transverse impression vague; basal fovez deep but not sharply defined, very dis- tinctly prolonged to middle, less distinctly nearly to apex; surface within and near fovez and across base transversely wrinkled-rugulose. Elytra subquadrate, virtually twice as wide as prothorax (in chalZybeus about I 7/10 times width prothorax) ; humeri rounded but not narrowed; sides faintly sinuate before independently narrowly rounded apices ; disk convex ; striz slightly impressed, faintly punc- tulate, except 2 external striz each side, and all stri~ at apex, much deeper and impunctate; intervals broad and slightly convex except 8th and 9th, and all at apex, narrow and very convex ; 3rd finely 3-punctate. Metepisterna very elongate; inner wings full ; body not punctate below. Tibiz finely grooved along middle of outer edge; posterior tarsi strongly grooved each side and at middle above, grooves ap- proximate so joints seem bicarinate, 4th joint emarginate, outer lobe somewhat longer than inner; front tarsi also grooved both laterally and at middle (these tibia1 and tarsal characters about as in chalybeus). Length 7.6; width about 3.2 mm.
JAMAICA: holotype 8 (M. C. Z. no. 22011), unique, from main range Blue Mts., about 5,000 ft., Aug. 17, 1934 ; shak- en from a pile of dead? cut grass in a narrow road through forest at Portland Gap, Blue Mt. Forest Reserve. Allied to C. chalybeus Dej.
(Lesser Antilles to Brazil),
of which we have specimens, but with broader elytra, with 8th and 9th intervals differently modified. Agonum (s. s.) laetificum n. sp.
Moderately broad9 rather depressed ; head and prothorax



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19351 West Indian Carabidz 201
moderately shining black, faintly greenish, elytra shining greenish coppery with margins not different; lower surface and appendages piceous. Head just under 213 width pro- thorax ; eyes only moderately prominent; gene short and oblique; antennz average, apex of 8th joint reaching about to base prothorax; mentum tooth acute. Prothorax rather large, rounded, just over 113 wider than long; sides arcuate from base to apex, margins rather narrowly ex- planate ; posterior angles broadly rounded ; base and apex finely margined; disk only slightly convex, middle line fine, transverse impressions vague, basal fovee rather shallow, not sharply defined, only vaguely continued anteriorly, not punctate. Elytra moderately (about 215) wider than pro- thorax; humeri somewhat rounded but not narrowed ; sides arcuate-subparallel at middle, sinuate before the apices, which are subindependently rounded ; striz moderate, not less impressed at apex, impunetate; intervals flat or slight- ly convex, polished, 3rd 3-punctate, first puncture attached to 3rd, 2nd and 3rd punctures to 2nd strize. Metepisterna with outer edges a little more than v2 longer than anterior ; inner wings full. Tibiz not sulcate on outer edge; hind tarsi with 3 basal joints sulcate each side, 4th not sulcate, shallowly emarginate; front tarsi not sulcate, at least in 8. Length 7.7; width 3.0 mm.
HAITI: holotype 8 (M. C. Z. no. 22023), unique, from summit plateau Mt. Basil, 4,700 ft., Sept. 9; among loose dead leaves on the ground in a small patch of wet cloud forest.
Very similar to Agonum cupripenne Say, especially to the form with the disk of elytra not strongly coppery, and probably an actual relative of that species, but broader and more depressed, with pronotum scarcely metallic, and with- out the conspicuous, bright green, strongly alutaceous mar- gin of elytra of cupripenne.
Chlaenius jamaicae n. sp.
Form average; head and prothorax rather shining dark bluish or green, elytra dark dull bluish; lower surface in- cluding epipleur~ dark; appendages testaceous, antennz a little brownish. Head v4 width prothorax ; eyes prominent ; front very finely punctulate; clypeus and labrum slightly



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202 Psyche [December
emarginate; antennae with 3rd joint equal to 4th, 7th about opposite base prothorax. Prothorax subcordate, 1/5 or wider than long, moderately narrowed in front and behind; base truncate, Vå wider than apex; sides arcuate through most of length, briefly and not strongly sinuate basally, narrowly margined, lacking anterior seta as usual ; basal angles slightly obtuse, narrowly blunted; disk convex, im- pressed and foveate as usual, rather closely punctate and slightly alutaceous across base, sparsely punctate and shin- ing anteriorly, with narrow impunctate area each side near middle. Elytra about 2/5 wider than prothorax, convex, very slightly narrowed basally; humeri rounded but not obliterated, marginal line arcuate; striae rather fine, faintly punctulate; intervals nearly flat, very finely granulate- opaque, pubescence shorter and less obvious than usual. Prosternum margined at apex ; metepisterna about % long- er than wide; sides of body below and entire abdomen rather finely punctate-pubescent. Femora and tarsi not densely pubescent above; 8 with front femora not modi- fied, middle tibiae not pubescent at apex. Length 11-12.5;
width 4.1-4.8 mm.
JAMAICA: holotype 8
(M. C. Z. no. 22024) and 8 para-
types from Ocho Rios, Aug. 20-24; 1 paratype each from Rio Cobre, 5 mi. above Spanishtown, Aug. 29 and between Kingston and Spanishtown, Aug. 29; all taken under cover beside ponds, swamps, or rivers.
Very close indeed to Chlsenius viridicollis Reiche and C. obscuripennis Chev., both of Mexico and both represented by small series in our collection (the two may be identical), but differing in the conspicuously shorter pubescence of elytra.
Stenocrepis (s. s.) palustris n. sp.
Subparallel, moderately broad and convex; black, mod- erately shining, only slightly aeneous with margins of elytra scarcely more so; lower surface and legs piceous; tarsi, palpi, and bases of antennae more or less rufescent. Head a little over y2 width prothorax, but appearing narrower; formed as in 12-striatus (Chev.) and similarly finely, close- ly subpunctulate ; mentum tooth short, wide, blunt. Pro-
thorax a trifle over 2/5 wider than long, moderately strong-



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19351 West Indian Carabidse 203
ly narrowed in front, a trifle behind; base % or more wider than apex ; sides nearly evenly arcuate throughout ; basal angles about right or a trifle obtuse; disk moderately convex, not punctate; middle line very fine, other impres- sions extremely vague or absent. Elytra only slightly (about 1/6) wider than prothorax, very convex; humeri distinct, denticulate ; sides straight, parallel or faintly diverging posteriorly for most of length, subapical sinua- tion and apices as in 12-striatus; each elytron with 6 mod- erately deep, smooth, straight striae as in 12-strmtus, 7th stria extremely faint; scutellar stria almost obsolete; inter- vals slightly convex, 3rd without dorsal punctures. Metepisterna and sides of metasternum very coarsely, first ventral coarsely, 2nd ventral and mesepisterna anteriorly much less distinctly punctate. Front tarsi 8 about as in 12-striatus, with 2nd joint about as long as wide. Length 7.5-8.5 ; width 3.1-3.6 mm.
HAITI: holotype 8 (M. C. Z. no. 22025) and 19 para- types from swamps north of Dessalines, Sept. 11; 3 para- types from Trou Caiman, Nov. 15-20; taken under cover in and beside swamps.
Resembles S. duodecimstriatus (Chev.), to which it runs in Chaudoir's monograph (Ann. Soc. Ent. France (6) 2, 1882, pp. 485- ), but relatively a little broader especially posteriorly, with shorter prothorax and more convex elytra, and with weaker seneous luster, characters which enabled me to distinguish the new species from 12-striatus even in the field. The two occurred together at Trou Caiman. The 3rd interval of each elytron is 2-punctate in 12-striatus, impunctate in palustris.
Slelenophorus flavilabris cubanus n. subsp. S. flavilabris Darl. 1934, Psyche 41, 105 (not Dej.) Very similar to true flavilabris Dej., of which I now have 171 specimens from Haiti and Jamaica (type locality: "Antilles"). Similar in variable, usually greenish or bluish, metallic color, highly polished surface, and im- punctate base and fovese of prothorax. Differs conspicu- ously in having the legs entirely testaceous; the femora are dark in true flavilabris.
The Cuban subspecies averages a
little smaller, but not sufficiently to be significant if it were



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204 Psyche [December
not for the difference in color of legs. Length 6.5-8.0; width 2.6-3.3 mm.
CUBA: holotype 8 (M. C. Z. no. 22026) and 35 paratypes from Soledad, near Cienfuegos, various dates and collectors. Bradycellus (Stenocellus) selleanus n. sp. Form about average for genus ; black, very polished, mar- gins of elytra (especially apically) translucent testaceous ; suture not pale; palpi, legs, base of mandibles, and base of antennae testaceous, apex of mandibles and antennae except basally darker. Head as usual in Stenocellus, about % width prothorax, impressions deep; antennae rather stout, 9th joint reaching- about to base prothorax; mentum tooth acute. Prothorax subcordate, about 14 wider than long; base about width apex; sides arcuate anteriorly, briefly but usually rather strongly sinuate before posterior angles ; latter briefly right or slightly obtuse, more or less strongly denti- form; disk as usual; basal foveae shallow, rather coarsely punctate. Elytra normal for genus, about 2/5 wider than prothorax, not very elongate; no scutellar stria; other striae entire, impunctate ; intervals slightly convex, polished, 3rd with single puncture on inner edge behind middle. Male front tarsi moderately dilated, biseriately squammu- lose ; middle tarsi slender, inconspicuously squammulose. Length 3.5-4.2 ; width 1.4-1.6 mm.
HAITI: holotype 8 (M. C. Z. no. 22027) and 31 paratypes from La Visite and vicinity, La Selle Range, 5,000-7,000 ft., Sept. 16-23; 3 paratypes from Kenskoff, a few miles north of type locality, about 5,000 ft., Nov. 8; common under ground cover in cloud forest and damp ravines. Comparable only with Bradycellus circumdcbtus Bates of Mexico (we have a small series from Biologia material), but smaller, much more polished, with sides of prothorax more briefly but much more strongly sinuate before base. Pogonodaptus rostratus n. sp.
Form of P. mexicanus (Bates) (piceus Horn) ; polished piceous, more or less rufescent, margins of elytra (espe- cially apically) testaceous ; legs testaceous with dark knees ; antennae brownish, testaceous at base. Head of 8 very



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19351 West Indian Carabidas 205
large, barely narrower than prothorax but appearing at least as wide; genae subprominent, subtuberculate; man- dibles long, not strongly arcuate ; clypeus emarginate, sub- tuberculate each side; labrum deeply emarginate; frontal sulci very deep, much closer together (where most approxi- mate) than to eyes; head in s about 6/7 width prothorax, with mandibles a little less prominent, gense short and not prominent, frontal sulci much less aproximate. Prothorax broadly cordate, or slightly more wider than long, much narrowed basally; base 1/10 to 1/5 narrower than apex; sides broadly rounded except very briefly sinuate before posterior angles ; latter would be rounded except for small, obtuse prominence; disk about as in mexicanus except more depressed and roughened across base, with narrower posterior margin, and with anterior transverse impression more distinct, lying behind the (interrupted) anterior mar- ginal line. Elytra about as in mexicanus. Measurements: large 8 8 by 2.5 ; small s 7.2 by 2.3 mm. HAITI: holotype 8 (M. C. Z. no. 22028) and 40 paratypes from swamps north of Dessalines, Sept. 11; in loose soil and piles of trash near the water.
As compared with mexicanus, of which we have a fine series from Texas (Brownsville) , Mexico (Vera Cruz) , and Colombia (near Santa Marta), the 8 of rostratus has more prominent mandibles and gene and much more ap- proximate frontal sulci (nowhere closer to each other than to the eyes in mexicanus). The thorax of the new species differs also, as noted in the description. Llebia haitiana n. sp.
Typical Lebia form ; weakly shining ; black, side margins of prothorax pale translucent testaceous ; elytra testaceous with basal blotch (between 4th striae), broad, slightly irregular post-median fascia (extending laterally to the marginal striae), a broad sutural connecting stripe (con- stricted nearly to first striae at basal 1/3) between the basal and post-median marks, a very narrow apical sutural line, and a small spot near the outer apical angle of each elytron black or dark brown; lower surface black; legs testaceous; palpi brown with pale tips. Head about 4/5 width prothorax; eyes very prominent; front longitudinally



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206 Psyche [December
rugulose; mentum tooth short, broadly rounded'; palpi not thickened. Prothorax transversely quadrate, about 2/3 wider than long, with anterior angles rounded, posterior bluntly right, preceded by a slight sinuation of the sides; explanate margins rather broad ; disk moderately convex, irregularly (transversely at middle, sublongitudinally at sides) rugulose. Elytra wide, about 5/6 wider than pro- thorax, moderately convex; striae rather deep, impunctate, except 3rd with usual 2 dorsal punctures on each elytron; intervals somewhat convex, alutaceous. Posterior tarsi with 4th joint bilobed; claws each with 4 rather long teeth below.
Length 5.3-5.7; width 2.6-2.8 mm.
HAITI: holotype 8 (M. C. Z. no. 22029) from vicinity of Port-au-Prince, Oct. 6; 1 8 paratype from Poste Terre Rouge, 2,000 ft., Oct. 5; 1 8 paratype from Port-au-Prince, Jan. 1925, G. N. Wolcott, from Andre Audant. My two specimens were taken by sweeping vines along overgrown roadsides.
The pattern of the elytra resembles, at first glance, that of Lebia, fuscata Dej., but closer examination shows that haitia-na lacks the dark marginal streak which, in fmcata, extends forward from the post-median fascia on each side, while fuscata has no separate posterior-lateral spot. The two species are very differently sculptured. In Chaudoir's key, haitiana runs to the analis group (Bull. Soc. Nat. Moscow 44, 1871, part 1, p. 23), but neither analis nor any of the other species listed has the thorax uniform black (except for the margins) as in haitha, nor does the pat- tern of the elytra of the latter coincide exactly with any of them.
Lebia nigrita n. sp.
Lebia-like, but with very convex elytra; black, weakly shining, appendages and explanate margins of prothorax brownish. Head, 6/7 width prothorax ; eyes prominent ; front alutaceous, not rugulose, with punctiform median impression ; mentum tooth short, broadly rounded ; palpi rather short but not much incrassate. Prothorax about 3/5 wider than long, transversely quadrate; anterior angles rounded, posterior right, with sides before them straight; explanate margins moderate ; disk moderately convex,



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1935 ] West Indian Carabidse 207
alutaceous, faintly transversely strigulose. Elytra more than 4/5 wider than prothorax, very convex but nearly normal in outline from above; striae deep, impunctate, except 3rd on each elytron with 2 unusually coarse dorsal punctures ; intervals moderately convex, alutaceous. Hind tarsus with 4th joint bilobed; each claw with 3 rather long teeth on inner edge.
Length 3.6 ; width 1.9 mm.
HAITI: holotype 8 (M. C. Z. no. 22030), unique, from hills south of Port-au-Prince (edge of Massif de la Selle), about 2,000 ft., Oct. 2; taken by sweeping path-side vines and weeds.
In Chaudoir's key this runs to scitula Chd. and allies (Z. c., p. 24), but is very much smaller than any of the species listed. It is probably actually more closely related to some of the very convex continental species such as hilaris Chd. and gramria Putz., but the uniform coloration and deep striae distinguish it.
Lebia gibba n. sp.
Of the same small, convex form as the preceding; rather shining blue, lower surface and appendages black. Head
about 5/6 width prothorax; eyes a little less prominent than in nigrita; front lightly alutaceous, not strigulose, with median puncture ; mentum tooth short and broadly rounded; palpi rather short but not much incrassate. Pro- thorax transversely subquadrate, 2/3 wider than long; anterior angles rounded; posterior angles almost right, sides before them straight but converging posteriorly ; sides of base slightly oblique toward rear, so that angles are directed slightly backward ; explanate margins moderate ; disk very convex, shining, neither punctate nor alutaceous. Elytra about % wider than prothorax, very convex, but with nearly normal marginal outline ; striae rather shallow, impunctate except for 2 rather small dorsal punctures on 3rd on each elytron; intervals slightly convex, rather shin- ing, only very faintly alutaceous. Hind tarsi with 4th joint bilobed; each claw with 3 teeth. Length about 3.3; width 1.7 mm.
HAITI: holotype s (M. C. Z. no. 22031), unique, from Hinche, Aug. 30, 1930, H. L. Dozier collector, from Mr. Andre Audant.




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208 Psyche [December
Much like the prece,ding (nigrita) in form, but differ- ently colored, more shining, with somewhat shallower ely- tral striae, and with somewhat differently formed prothorax, with much more convex disk.
Phloeoxena montana n. sp.
Form as in normal Phloeoxena except elytra more oval and much more convex; head and prothorax moderately shining, elytra duller; brownish piceous above and below, epipleurae of elytra and last ventral segment pale; femora testaceous, tibiae a little browner; palpi and antennae slight- ly brownish testaceous. Head about 85/100 width prothorax, normal for Phlceoxena except eyes a trifle less prominent than usual; front alutaceous, with 2 anterior and single median impressions weak ; antennae rather stout, 8th joint reaching about to base prothorax ; mentum tooth rounded at apex. Prothorax 1/6 or 1/7 wider than long, narrowed behind; base truncate, a little narrower than apex; sides weakly arcuate anteriorly, straight and converging in about posterior s; posterior angles obtuse, somewhat rounded; anterior angles somewhat produced, narrowly rounded ; lateral margins narrow, without setae ; base margined, apex indistinctly so; disk convex, finely alutaceous; middle line well impressed except at ends, transverse impressions and basal foveae almost obsolete. Elytra, broadly oval, with re- duced, broadly rounded humeri ; margins, marginal setae, and apical sinuato-truncation normal ; disk convex, rather strongly alutaceous ; striae faint, broad, and vaguely de- fined, about as in schwarzi Darl.; each 3rd interval with 2 dorsal punctures about 2/5 and 1/10-from apex, usual anterior puncture lacking, extreme basal foveolate pune- ture (contiguous to margin) present. Metepisterna very short, outer edges no longer than anterior; inner wings atrophied. Body below alutaceous, sides of abdomen some- what wrinkled. Hind tarsi normal, i. e. with 4th joint a little shorter than but otherwise similar to third; claws slender and rather elongate, each with 3 teeth on inner edge, outer tooth scarcely outside of middle, so apex of claw is long, slender, and simple. Male front tarsi very weakly dilated, each with 3 joints biseriately squammulose ; middle



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19-36] West 1ndW.n Carabidse 209
tarsi not modified; last ventral with 1 ( 8 ) or 2 ( 9 1 sek each side. Length 3.8-5.0; width 1.6-2.0 mm. HAITI: holotype 8 (M. C. Z; no. 22032) and 11 para- types from Mt. La Hotte, on and near summit (probably not below 7,000 ft.) , Oct. 16-17 ; in leaf mold, moss, and rotting wood in damp forest.
Similar in color to P. schwarzi Darl., but differing from it, and I think from all other known Phlceoxena, by the loss of wings and consequent modification of elytra. Apenes apterus n. sp.
Form as usual in Apenes s. s., Sznmta group ; moderately shining purplish bronze, humeri with poorly defined brown spots, outer margin of elytra vaguely and irregularly brownish ; lower surface piceous ; appendages brown. Head large, about 6/7 width prothorax, but with eyes less promi- nent than usual ; front finely alutaceous, finely and sparsely punctulate, not strigdose; antennas rather short, middle joints scarcely longer than wide. Prothorax % wider than long, subcordate; sides arcuate anteriorly, virtually straight and converging or slightly sinuate before posterior angles; latter either almost obliterated or very minutely subpromi- nent; disk finely alutaceous, without evident punctulation, and with faint, sparse, transverse wrinkles. Elytra, I/^ wider than prothorax; humeri about as in winged species; strise moderate, not punctate; scutellar stria present but rather variable ; intervals slightly convex, lightly alutaceous, 3rd with 2 conspicuous punctures, at about 1/3 and 2/3 of length. Metepisterna with outer edges about I/^ longer than anterior, shorter than in winged species; inner wings atrophied. Last joint labial palpus 8 not nearly so dilated as usual in genus, scarcely wider than in 9, with apical much shorter than inner margin ; last ventral with 2 setae each side in both sexes. Length (to apex elytra) 6.6-7.0; width 2.5 mm. or slightly more.
JAMAICA : holotype & {M. C. 2. no. 22033), and 2 Q s paratypes from main range Blue Mta., 5,000-7,000 ft., Aug. 17-19; under stones along the trail in the forest. Differs from the other West Indian species of the Sznuata group (delicata Dad. etc.) in lacking a subapical pale fascia, and in the atrophy of the wings.



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210 Psyche [December
Apenes ovalis n. sp.
Broad, with elytra more oval and more convex than usual; shining piceous without pale markings above; ap- pendages and elytral epipleurse reddish testaceous. Head large, a trifle over %. width prothorax; eyes but little prominent; front shining, very faintly alutaceous and punctulate, not strigose ; antennae rather short, middle joints a little longer than wide. Prothorax broad, 2/5 to % wider than long; sides broadly rounded, almost evenly rounded into base except for vague lobe just behind pos- terior seta each side; base also vaguely lobed; disk mod- erately convex, shining, very faintly alutaceous, faintly transversely wrinkled near middle. Elytra broadly oval, 1/^ wider than prothorax ; striae moderate, impunctate ; scutellar stria rather short; intervals flat or faintly con- vex, polished, 3rd with 2 strong punctures. Metepisterna with outer edge about 1/^ longer than anterior, shorter than in winged species ; inner wings reduced to minute vestiges. Male with apical joint labial palpus much less dilated than usual, apex scarcely more than 14 long as inner edge, scarcely wider than in 9 ; 8 with 1, 9 with 2 setae each side last ventral. Length (to apex elytra) 8.0-9.5 ; width 3.4-3.9 mm.
HAITI: holotype 8 (M. C. Z. no. 22034) and 55 paratypes from La Visite and vicinity, La Selle Range, 5,000-7,000 ft., Sept. 16-23; in loose leaf mold and under other cover in wet cloud forest.
The wide, oval, convex elytra, and shining, unspotted surface give this Apenes a unique appearance which dis- tinguishes it easily from the preceding (aptera), which is the only other species of the genus known to me from the West Indies in which the wings have atrophied. Eucserus haitianus n. sp.
Stout, moderately convex ; black ; head and prothorax alutaceous, elytra shining, subiridescent; antennae with 6 proximal joints, brownish, 7th more or less bicolored, 4 distal whitish; palpi, labrum, and legs brown, latter some- times more or less testaceous. Head as in insularis Darl., with middle antenna1 joints about 3 times long as wide. Prothora,~ broadly cordate, slightly to distinctly more than



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19351 West Indian Carabidse 211
y^ wider than long; sides briefly but distinctly sinuate be- fore minute, somewhat obtuse basal angles; base lobed at middle, slightly or (usually) distinctly sinuate each side. Elytra with rather fine, entire, impunctate striae ; intervals barely convex; 3rd stria with fine setigerous puncture about 1/5 from base, 2nd with similar punctures near middle and a little before apex. Inner wings reduced as usual. Male with front tarsi slender, inconspicuously squammulose ; 8 with 1, $ with 2 setae each side last ventral. Length (to apex elytra) 3.6-4.6; width 1.6-2.1 mm.
HAITI: holotype $ (M. C. Z. no. 22035) and 17 para- types, mostly $ $, from Etang Lachaux, Oct. 26-27 ; taken by raking leaves from shady places on the bank into the water.
This species is closely allied to the Cuban insularis Darl. (Psyche 41, 1934, p. 120), of which I took 20 additional specimens at the type locality
(Soledad), Aug. 2-12, 1934,
but haitiw differs constantly in the brown labrum (testa- ceous in insularis) and in having a wider prothorax with posterior angles better defined and sides of base sinuate. Pentagonica flavipes picipes n. subsp.
Structurally the same as typical P. flavipes (Lee.) of North America and Cuba (we have 33 specimens from Soledad, near Cienfuegos) and similarly uniform piceous black above, with fine elytral strise, but with legs piceous or dark brown instead of testaceous or very pale brown and with labrum and base of antennae somewhat darker also. The latter, however, are rather variable in true flavipes. The difference in color of legs is conspicuous even in the field, to the naked eye.
GREATER ANTILLES except CUBA: Jamaica: holotype $ (M. C. Z. no. 22036) from Kingston, Aug. 27; 1 paratype from Ocho Rios, Aug. 24; 5 paratypes from Port Antonio, March, E. A. Wight collector. Haiti: 1, not a type, from Miragoane, Oct. 30. Puerto Rico: 1, not a type, from S. T. Danf orth.
Galerita montana n. sp.
Rather stout and convex ; elytra costate ; black or piceous, appendages dark, with 7 outer joints of antennae dull



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212 Psyche [De,cember
rufous. Head about % width prothorax; eyes scarcely prominent; gense rounded, nearly wide as eyes; front and vertex rugosely punctate except for small smooth area at middle anteriorly. Prothorax between subquadrate and subcordate, as wide to 1/10 wider than long, somewhat narrowed basally ; sides strongly arcuate anteriorly, rather broadly sinuate before right or slightly acute, but nar- rowly rounded, posterior angles ; disk moderately convex, rugosely punctate; middle line fine, transverse impressions not sharply defined ; basal fovese small and deep. Elytra rather broadly oval, with broadly rounded, somewhat nar- rowed humeri; sides just behind humeri more or less con- vex in outline, rarely almost straight, never subsinuate; apices strongly truncate, each subemarginate, with outer apical angles well defined, though obtuse ; disk rather con- vex; each elytron with 9 moderate primary costse plus a rather elongate scutellar costa (comparable to a scutellar stria in position) ; each primary intercostal interval with 2 much finer cost=, except only 1 secondary costa at suture and none, or only faint traces, in marginal interval; sur- face between costse finely transversely rugulose. Mete- pisterna about Vy, longer than wide; inner wings vestigial. Length 15.5-18 (to apex elytra, 14-16) ; width 5.2-6.2 mm. HAITI: holotype 8 (M. C. Z. no. 22037) and 14 para- types from vicinity of La Visite, La Selle Range, 5,000- 6,000 ft., Sept. 16-23 ; under stones in damp places on high slopes, but not on the plateau (except just at the edge) and not in cloud forest.
Somewhat similar to Galerita nigra Chev. of Mexico, but with outer apical angle of elytron much better defined and differing slightly in other ways. Probably more close- ly related to the Haitian striata Klug, of which we have a series from various localities below 5,000 ft., but larger and broader, with relatively slightly broader prothorax, and with elytra more evenly oval, without the post-humeral subsinuation of striata.
Pseudaptinus marginicollis f umipes n. subsp. Structurally the same as typical Cuban marginicollis Darl. {Psyche 41, 1934, p. 126) except that the prothorax is slightly broader anteriorly in the new subspecies; color



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19351 West Indian Carabidse 213
also similar except the legs either entirely (7 specimens)
or with only the tibiae and apices of the femora (2 speci- mens) brown or blackish, while in typical rnarginicollis, of which I now have 28 specimens from the type locality (Soledad, near Cienfuegos, Cuba), the legs are always en- tirely pale testaceous.
HAITI : holotype (M. C. Z. no. 22038)
(with legs entirely
dark) and 4 paratypes (3, legs entirely dark; 1, with only tibiae and apices of femora dark) from Ennery, Sept. 6-11; taken from piles of old flood debris beside the river; addi- tional paratypes : 1, swamps north of Dessalines, Sept. 11 ; , 1, Miragoane, Oct. 30; 1, Camp Perrin, Oct. 9; 1, Etang Lachaux, Oct. 26:
Zuphium haitianum n. sp.
Normal Zuphium form; piceous, a little more brownish on elytra, suture and medio-basal area of elytra and small vague sublateral stripes of prothorax usually faintly red- dish ; abdomen (irregularly) and legs testaceous ; mouth parts including labrum, and antennae rufo-testaceous. Head slightly (about 1/10) narrower than prothorax, shaped as in cubanurn; front finely, not closely punctulate, finely alutaceous, more distinctly so on vertex; antennae with joints 5 to 8 twice or slightly more times long as wide. Prothorax barely (not over 1/10) wider than long, sides of apex slightly less oblique than in cubanurn (i. e., prothorax more square shouldered) ; sides broadly arcuate anteriorly, strongly sinuate before the right or slightly acute posterior angles ; disk moderately convex ; middle line sharp, anterior transverse impression obsolete, posterior rather vaguely impressed; basal foveae deep, rounded; whole pronoturn finely, closely punctate and finely alutaceous. Elytra about 7/10 wider than prothorax, a trifle shorter and wider pos- teriorly than in cubanurn; striae moderate ; intervals slightly convex, rather finely roughened and alutaceous. Length (to apex elytra) 4.0-4.7; width 1.4-1.6 mm. HAITI: holotype (M. C. Z. no. 22039) and 9 paratypes from Etang Lachaux, Oct. 26-27; taken by raking dead leaves from shady places into the water. Duller and much more finely punctate on head and pro- thorax than cubanurn Liebke, of which we have 29 speci-



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214 Psyche
[December
mens from Soledad, near Cienfuegos, Cuba. In Liebke's
key to the American Zuphium
(Revista Ent. 3, 1933, pp.
461- ) haitianum runs to columbianulm Chd., but the latter (which I have not seen)
is evidently a pale species with
only the head dark, and color is a rather constant character in this genus.
Pse~d~mo~rpha caribbeana n. sp.
Slender, parallel, very convex (even more so than in ang~sta~ta Horn) ; castaneous, lower surface (except abdo- men) and appendages more rufescent ; whole upper surface finely alutaceous under high power, merely silky under lower power, moderately shining. Head 7/10 width pro- thorax; front with transverse row of moderate, setigerous punctures between eyes, otherwise very finely, sparsely punctate ; pre-ocular lobes moderate, more explanate than in angustata; clypeal suture very fine; antennae slender, reaching just beyond anterior coxse. Prothorax 2/5 wider than long; apex truncate except anterior angles slightly prominent; sides broadly, almost evenly arcuate, with fine margins ; basal angles rounded ; apex margined, base not ; disk finely, sparsely punctate and pubescent; middle line excessively fine ; base not impressed, baso-lateral impres- sions completely absent. Elytra probably no wider than prothorax (slightly distorted by pin), elongate; sides parallel ; apices rounded-truncate ; suture not elevated ; each elytron with 9 rows (beside the marginal fringe) of setig- erous punctures, no special rows coarser than others; punc- tures so widely spaced in the rows that at first glance the elytra appear punctured at random ; surface otherwise im- punctate. Length 7.2 ; width 2.7 mm.
HAITI: holotype, probably $ (M. C. Z. no. 22040), unique, from Port-au-Prince, May, 1927, G. N. Wolcott collector, received from Mr. Andre Audant.
This is the first species of subfamily Pseudomorphinse to be known from the West Indies. It is a very distinct species. In Notman's key to the species of Pseudomorpha (Proc. United States Nat. Mus. 67, 1925, Art. 14, pp. 14-15) the Haitian species runs to confusa Notman, but, as com- pared with the description of confusa, caribbeana is small- er (confusa, 10.25 mm.), much more convex, with relative-



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19351 West Indian Carabidas 215
ly wider head and narrower prothorax, and differs some- what in other ways. Notman described confuse from a unique 2 from the British Museum supposed to come from Australia ; since Pseudomorpha is otherwise entirely con- fined to a small part of America, however, the Australian record is probably erroneous.




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