H. H. Ross.
Lectotypes of North American Caddis Flies in the Museum of Comparative Zoology.
Psyche 45(1):1-61, 1938.

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PSYCHE
VOL. XLV MARCH, 1938 No. 1
LECTOTYPES OF NORTH AMERICAN CADDIS FLIES IN THE MUSEUM OF COMPARATIVE ZOOLOGY1
Illinois State Natural History ' Survey, Urbana, Illinois
Few species of caddis flies described by Hagen and Banks have had lectotypes designated for them. In a few cases the type series of one species contains representatives of more than one species, a condition which has led to confusion of names due to different interpretations made by different authors. The only way to obviate further repetition of this is to place on a single type basis (by lectotype designations) all those species described from a series of eotypes. This paper sets lectotypes for 229 species, all the lectotypes being in the collection of the Museum of Comparative Zoology. The method followed in selecting the single type specimens has taken into account very little work done by other au- thors, since a large number of the species represent segre- gations seldom recognized by others. Furthermore both Hagen and Banks used a labelling system which left no doubt as to which specimen they considered the real type. Hagen placed his label on only one specimen and marked it with an asterisk if it were a type or plesiotype in the sense of present usage. Other specimens in the series were simply placed in 1A grant from the travel fund of the Illinois State Natural History Survey, Urbana, Illinois, made it possible for me to visit the Museum of Comparative Zoology and make these studies. I wish to acknowl- edge my gratitude and appreciation to the Survey for this grant. Pachc 45:l-61 11938). http //psychr enlclub.me/-IS/45-001 him)



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2 Psyche
[March
a row after the first labelled specimen; undoubtedly some specimens which are not actually types were put in these series at the same time or subsequently, either by Hagen himself or inadvertantly by others. In selecting lectotypes of Hagen's species the specimen labelled by Hagen has been taken in each case. Lectotypes have been designated for all the species from North America described by Hagen unless they have been previously set in literature, even if only rep- resented in the collection by a single individual. This is done because of the possibility of specimens appearing in other collections which might erroneously be considered as types.
There is a possibility that some specimens designated as lectoallotypes have not been correctly associated with the lectotypes, that is, do not represent the opposite sex of the same species as the lectotypes.
In a very high proportion of
the North American caddis fly species it is impossible, with our present studies, to separate the females of closely re- lated forms, so that we have to rely to an inordinate extent on collection data as a basis for association. All the specimens mentioned as "allotype" represent des- ignations made for the first time in this paper. In the Banks collection the specimens listed without de- finite collector from the eastern states were taken by Banks himself, except in a few cases.
Banks' species have his own label on the first specimen and simply a "TYPE" label on the others. The first specimen with the label has been taken as the lectotype, a procedure which Mr. Banks has requested and which seems perfectly logical.
In rare cases when these first specimens have been females and the cotype series contained males, an exception to the foregoing procedure has been instituted and a male selected as the lectotype.
No attempt has been made to analyze any of the cotype series except the lectotypes, the remainder automatically becoming paratypes. Such a study would have little signifi- cance since the paratypes will have only historic and minor taxonomic interest. Furthermore so many of the specimens have been sent in exchanges, etc., to institutions in widely



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19381 North America,n Caddie Flies 3
scattered countries that it has been impossible for me to gather the necessary information.
The species treated are listed within each family alpha- betically according first to genus and then to species. Many of the species have been transferred from genus to genus several times. In attempting to find some method which would be easy to follow and at the same time show the generic placement, I am using the following double entry when a change has been made. If a species is placed in a genus other than the one in which it was described, it is listed both under the genus in which it was originally de- scribed and under the genus in which I am placing it. For example, Hydropsyche sordida Hagen is now placed in the genus Cheumatopsyche. Therefore, it is listed under both Hydropsyche sordida in the h's and under Cheumatopsyche sordida. Under the former, note is made of the genus in which it is placed.
Professor Nathan Banks, Curator of insects at the Mu- seum of Comparative Zoology, and Professor F. M. Carpenter of the Division of Biology, Harvard University, have been of inestimable help during the course of this work in giving information and advice, in placing at my disposal study facil- ities and equipment, and in the many courtesies accorded me during my visit to the Museum of Comparative Zoology. Dr. C. 0. Mohr, of the Illinois State Natural History Sur- vey, has made most of the drawings for this paper, and I wish to express my gratitude for this.
Family RHYACOPHILID2E
Agapetus malleatus Banks, 1914, p. 202, fig. 57. Lectotype, male.-Los Angeles County, California, San Gabriel Mts., 3000 ft., June 17, 1907, F. Grinnell, Jr. No. 11723. Lectoallotype, f emale.-Same data. The genitalia of the lectotype are shown in fig. 1. Baerea? maculata Hagen.-see Protoptila maculata ( Hagen) Glossosoma nigrior Banks.-see Mystrophora nigrior (Banks)
Glossosoma parvulum Banks, 1904a, p. 108, fig. 13. Lectotype, male.-Pecos, New Mexico, August 10, at



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4 Psyche [March
light. No. 11748. Lectoallotype, female.-Same data, but August 13.
The genitalia of a homotype are shown in fig. 2. Mystrophora lividum (Hagen), 1861, p. 295 (Tinodes) . Lectotype, female.-St. Lawrence River, Canada, Osten Sacken. No. 11081.
Milne has determined the male of this species as that illustrated by Betten as Mystrophora sp. (1934, pi. 9, figs. 15-17).
Mystrophora nigrior (Banks), 1911, p. 355, fig. 23. ( Glossosoma)
Lectotype, male.-Black Mountain, North Carolina, north fork Swannanoa River, May. No. 11745. Lectoallotype, f emale.-Same data.
At present there appear to be two valid species in this genus, the two illustrated by Betten (1934) as americana and Mystrophora sp. No good characters have been dis- covered as yet which separate the females of the two species, so that the exact status of the names americana Banks and Uvidum Hagen is problematic. It seems best at present to consider as nigrior the species treated as americana by most authors and illustrated by Betten (1934, pi. 8, figs. 4-6 and pi. 9, figs. 1-14), and to consider americana as an unidentified species.
Protoptila maculata (Hagen), 1861, p. 296. (Baerea?) Lectotype, male.-St. Lawrence River, Canada, 1859, Sacken. No. 11093. Lectoallotype, female.-Same data. This species has been illustrated by both Banks and Betten (1934).
Rhyacophila acropedes Banks, 1914, p. 201, fig. 39. Lectotype, male.-Deer creek, Provo Canon, Utah, Au- gust 21, Spalding. No. 11741.
This species is closely allied to coloradensis Banks, but differs in details of the male genitalia, fig. 6. Rhyacophila atrata Banks, 1911, p. 351.
Lectotype, male.-Black Mountain, North Carolina, north fork Swannanoa River, May. No. 11739. Lectoallotype, f emale.-Same data.
The genitalia of the type are shown in fig. 3.



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19381 North American Caddis Flies 5
Rhyacophila brunnea Banks, 1911, p. 252. Leetotype, f emale.-Beulah, New Mexico, July 16. No.
11735.
There is a series of males and females in the M. C. 2. from Cultus Lake, B. C., the females of which appear exactly like the lectotype of brunnea in color and external genitalia. A
male of this series has been selected as the allotype. It be- longs to the acropedes group, fig. 4, but is readily distin- guished by details of the genitalia.
Allotype, male.-Cultus Lake, British Columbia : May 18, 1933, W. E. Ricker.
Rhyacophila Carolina Banks, 1911, p. 353, fig. 31. Leetotype, male.-Black Mountain, North Carolina, north fork Swannanoa River, May. No. 11727. Lectoallotype, female.-Same data.
In addition to those in the original description, illustra- tions of the genitalia have been given by Betten (1934, pi. 5, figs. 4-7).
Rhyacophila coloradensis Banks, 19 05b, p. 10. Leetotype, male.-Fort Collins, Colorado. No. 11728. This species is close to bifila Banks, but is readily distin- guished by details of the genitalia, fig. 5. Rhyacophila fairchildi Banks, 1930a, p. 130, figs. 4, 7. Leetotype, male.-Baddeck, Cape Breton Island, Nova Scotia, September 4, 1928. No. 16237. Lectoallotype, fe- male.-Same data.
The genitalia of the lectotype agree perfectly with the illustrations of glaberrima Ulmer and with the genitalia of the type of andrea Betten. There seems little doubt, there- fore,
that both fairchildi and andrea are synonyms of Ulmer's species, originally described from Georgia. Rhyacophila formosa Banks, 1911, p. 353. Leetotype, female.-Delaware Water Gap, New Jersey. No. 11078.
To date no male has been associated with this form, draw- ings of which are given by Milne (1936). Rhyacophila grandis Banks, 1911, p. 350, fig. 27. Leetotype, male.-Bon Accord, British Columbia, June 14. No. 11737. Lectoallotype, female.-Same data, but June 19.



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6 Psyche [March
The genitalia of the lectotype are well exposed without clearing, including the apex of the side tubes on the sedeagus. They are identical with those of the homotype illustrated in fig. 8.
Rhyacophila hyalinata Banks, 1905b, p. 10. Lectotype, male.-South West Colorado, July 23, 1899. No. 11738.
The genitalia of the lectotype have been cleared and com- pared critically with those of the specimen illustrated in fig. 7.
Rhyacophila luctuosa Banks, 1911, p. 351, fig. 24. Lectotype, male.-Woodworth Lake, Fulton County, New York, June 23, 1910, Alexander. No. 11740. Lectoallotype, f emale.-Same data.
The male genitalia are shown in fig. 9.
The species is
listed as a synonym of invaria Walker by Banks (1930b). Rhyacophila minora Banks, 1924, p. 444, fig. 37. Lectotype, male.-White Mountains, New Hampshire, Morrison. No. 14857. Lectoallotype, female.-Same data. This species was described and illustrated by Betten (1934, p. 134, pi. 7, figs. 10, 11) as Rhyacophila sp. 1. Rhyacophila nevadensis Banks, 1924, p. 443, fig.' 53. Lectotype, male.-Reno, Nevada, Morrison, 1878. No. 14855. Lectoallotype, f emale.-Same data. Rhyacophila nigrita Banks, 1907a, p. 132, fig. 16. Lectotype, female.-Black Mountain, North Carolina, June. No. 11742.
The male of this species has been figured in considerable detail by Betten (1934, pi. 7, figs. 1-5). The allotype will have to be designated' from one of his specimens. Rhyacophila rotunda Banks, 1924, p. 443, fig. 33. Lectotype, male.-Reno, Nevada, 1878, Morrison. No. 14856. Lectoallotype, f emale.-Same data. Rhyacophila torva Hagen, 1861, p. 296.
Lectot ype, male.-Washington, D. C., Sacken. No. 11078. The genitalia of this species have been illustrated by Banks (1907a, p. 132, fig. 7) and Betten (1934, pi. 7, figs. 6-9).




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19381 North American Caddis Flies 7
Tinodes lividum Hagen.-see Mystrophora lividum (Hagen) Family PHILOPOTAMIDX
Chimarrha aterrima Hagen, 1861, p. 297.
Lectotype, male.-Mus. Berol. Penn. No. 11098. The male genitalia have been illustrated by Betten (1934, pi. 16, figs. 6-9).
Chimarrha femoralis (Banks), 1911, p. 358. ( Womaldia) .
Lectotype, male.-Sacandaga River, Sport Island, New York, June 27,1910. No. 11520.
This species is a synonym of soda Hagen. Betten has
figured the genitalia under the latter name (1934, pi. 16, fig. 13).
Chimarrha plutonis (Banks), 1911, p. 358, fig. 34. (Wor- maldia) .
Lectotype, male.-Delaware Water Gap. No. 11519. The male genitalia of this species have been illustrated by Betten under the name lucia Betten (1934, pi. 16, figs. 10-12).
Chimarrha texana Banks, 1920, p. 360.
Lectotype, female.-San Antonio, Texas, August, Snow. No. 10914.
To date no reliable characters have been found for sepa- rating the females of this genus, so that for the present this species must be considered of doubtful identity. Dolophiliella gabriella Banks.-see Dolophilus gabriella (Banks).
Dolophilus breviatus Banks, 1914, p. 254, fig. 61. Lectotype, male.-Ithaca, New York, in Coy Glen, August. No. 11518.
The male genitalia of this type are broader than those of moestus, but the study of additional specimens indicates that this difference may be only individual variation. Dolophilus gabriella (Banks), 1930b, p. 230, fig. 14. (Dolo- philiella) .
Lectotype, male.-San Gabriel Mountains, California, June 29. No. 16326.
The most distinctive feature of this species is the more or



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8 Psyche
[March
less spatulate process of the eighth sternite. Dolophilus major Banks, 1914, p. 254, fig. 66. Lectotype, male.-Black Mountain, North Carolina, May. No. 11517.
This is the largest eastern species in the family. Philopotamus aequalis Banks, 1924, p. 450, fig. 48. Lectotype, male.-Tolland, Colorado, Dodds. No. 14853. Lectoallotype, f emale.-Same data.
This western species is readily distinguished by its dark color and the male genitalia.
Philopotamus americanus Banks, 1895, p. 316. Lectotype, male.-New York. No. 11512.
Both Banks and Betten have given illustrations showing a difference between this species and distinctus Walker. While these differences do occur, I have found all inter- gradations between them in a study of specimens from Michigan, New York, North Carolina and Maryland. All collections which I have seen containing specimens of both sexes are alike in having the females with minute, vestigial wings. This evidence indicates that only one species is involved in this material.
Wormaldia femoralis Banks.-see Chimarrha femoralis (Banks).
Wormaldia plutonis Banks.-see Chimarrha plutonis (Banks).
Family HYDROPTILID2E
Agraylea fraterna Banks, 1907b, p. 164.
Lectotype, male.-Falls Church, Virginia, May 1. No. 11591.
This species is a synonym of multipunctata Curtis. Allotrichia flavida Banks, 1907b, p. 164. Lectotype, female.-Fort Collins, Colorado, June 9. No. 11593.
No males have yet been associated with this form, so that its exact placement cannot be given.
Allotrichia maculata Banks.-see Hydroptila macvlata (Banks).




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19381 North American Caddis Flies 9
Hydroptila albicornis Hagen, 1861, p. 275. Lectotype, male.-St. Lawrence River, Canada, Osten Sack,en. No. 11105.
This species is easily identified by the sedeagus and clasp- ers, fig. 10. Hagen says the type is a female, but he was evidently misled by the retracted condition of the genitalia. Hydroptila maculata (Banks), 1904b, p. 116, 3 figs. (Allotrichia) .
Lectotype, male.-Falls Church, Virginia. No. 11595.
The unique male genitalia, fig. 11, are approached only by waubesiana Betten, but many differences separate the two. Hydroptila tarsalis Hagen.-see Polytrichia tarsalis (Hagen).
Hydroptila transversa Banks, 1907b, p. 163. Lectotype, male.-Washington, D. C., September, at light. No. 11592.
This is a synonym of maculatus (Banks).
Orthotrichia americana Banks, 1904b, p. 116, 1 fig. Lectotype, male.-Washington, D. C., August 13. No. 11598. Lectoallotype, female.-Same data. Illustrations of the cleared male genitalia have been given by Morton, 1905, under the name brachiata, which falls as a synonym of americana Banks. The species americana of authors will take the name cristata Morton. Orthotrichia nigritta Banks, 1907b, p. 163, figs. 1-3. Lectotype, male.-Austen, Texas, March 3, 1901. No. 11596.
This striking, black species from the southwest is different in many respects from the usual definition of Orthotrichia and should be placed in a new genus.
Metrichia new genus
Characteristics.-Ocelli present, close to eye. Tibia1 spur count, 1-3-4; spur on front tibia apical and small. Wings, fig. 14, narrowing to a pointed apex. Front wings with Sc very stocky, and all the other veins well developed; Ml-2 almost fused with Rs; Cu & Cu2 running close to the hind margin of the wing. Hind wing with Rl apparently reduced



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10 Psyche [March
to a short "cross-vein", and MI-2 fused for a short distance with Rs.
Genotype.-Orthotrichia nigritta Banks (by original designation).
This genus keys out with Stactobia but differs from it in the position of Cu & Cu2, the shortening of Rl, and in other characters.
Orthotrichia pallida Banks.-see Oxyethira pallida (Banks) Orthotrichia pictipes Banks.-see Stactobia pictipes (Banks)
Oxyethira dorsalis Banks, 1904d, p. 216, pi. 2, fig. 5. Lectotype, f emale.-Falls Church, Virginia, June 26, No. 11600.
That portion of the type series which corresponds to the original description, especially in having the characteristic white line down the dorsum, contains only female specimens. Until the females and males of more species in the genus have been associated nothing can be done to settle the specific identity of this species.
Oxyethira pallida (Banks), 1904d, p. 215, pi. 2, figs. 2, 7. Lectotype, male.-Washington, D. C. No. 11599. Lecto- allotype, f emale.-Same data.
This species is the same as viminalis Morton, which now becomes a synonym of pallida.
Polytrichia tarsalis ( Hagen) , 1861, p. 275. (Hydroptila)
Lectotype, male.-St. Lawrence River, Canada, Osten Sacken. No. 11104.
This species has been illustrated by Betten (1934, pi. 12, figs. 11-14) under the name confusa Morton. Morton's species is not this one, but according to original figures be- longs to another section of the genus.
Stactobia pictipes (Banks), 1911, p. 359. Lectotype, male.-Johnstown, New York, June 28, Hale's Creek. No. 11597. Lectoallotype, female.-Same data. The spur of the front tibiae put this species in the hetero- geneous Stactobia.
The male genitalia are illustrated in
fig. 12. They are strikingly different from any other ~earctic species.




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19381 North American Caddis Flies
Family POLYCENTROPODIDB
Cernotina pallida (Banks), 1904d, p. 214. (Cyrnus) Leetotype, male.-High Island, Maryland, June 17. No. 11539.
This interesting species belongs in the subfamily Psy- chomyiin~.
The male genitalia are very distinctive, fig. 18. The only females in the M. C. Z. under this name do not belong to this species, so that as yet an allotype cannot be designated.
Cyrnus fraternus Banks.-see Nyctiophylax fraternus (Banks)
Cyrnus pallidus Banks.-see Cernotina pallida (Banks) Plectrocnemia albipuncta Banks.-see Pol ycentropw a1 bi- punctw (Banks)
Plectrocnemia aureola Banks? 1930a, p. 130, figs. 2, 3, 5. Lectotype, male.-Baddeck? Cape Breton Island, Nova Scotia, July 209 1928. No. 16323. Leetoallotype, female.- Same data.
Illustrations of the male genitalia are given in fig. 13. Plectocnemia cinerea (Hagen), 1861, p. 293. (Polycen- tropus)
Le~totype~ male.-St. Lawrence River, Canada, 1859? Osten Sacken. No. 11039. Lectoallotgpe, f emale.-Same data.
The genitalia of this species have been illustrated by Betten (1934, pl. 24, figs. 1-8) under the specific name canadensis Banks.
Plectrocnernia flavicornis (Banks), 1907b, p. 162, fig. I. (Holocentropus )
Lectotgpe, male.-Washington? D. C. No. 11526. This is a synonym of cinerea (Hagen).
Plectrocnemia paNescens Banks, 1930b9 p. 231, fig. 3. Lectotype? male.-Put-in-Bay, Ohio, July 3? 1924, on Middle Bass Island, G. Townsend. No. 16322. Lectoallo- type, female.-Same data, but July 6, 1926. The genitalia of this specimen appear identical with those of cinerea (Hagen), but the color is lighter. It is my belief that the specimen is slightly teneral. In several species



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12 Psyche [March
of caddis flies I have taken large collections every specimen of which was uniformly teneral.
Holocentropus flavicornis Banks.-see Plectrocnemia f‰âÂav cormk (Banks)
Holocentropus interruptus Banks, 1914, p. 257? fig. 71. Le~totype~ male.-Hampton? New Hampshire? June 15, 1908. No. 11543.
This species was illustrated by Betten (1934? pl. 24, fig. 9). It has line priority over orotus Banks.
Holocentropus longus Banks, 1914, p. 258? figs. 65? 68. Leetotype? f emale.-Framingham? Massachusetts? June 4, 1904? C. A. Frost.
No. 11542.
The size and color of this specimen leaves little doubt but that it is the same as interrupt~s~ although sure diagnostic characters have not yet been discovered for the females of this genus.
Ho~~centropus orotus Banks? 1914? p. 257, fig. 69. Lectotype, male.-Clear Creek? Colorado. No. 11541. Lectoallotype? female.-Chimney Gulch, Boulder, Colorado, Oslar.
This species is a synonym of intermptus Banks, which has line priority.
Holocentropus placidus Banks.-see Phylocentropus placidus (Banks)
Neureclipsis parvulus Banks? 1907b? p. 163? figs. 2? 3. Le~totype~ male.-High Island? Maryland, June 17. No.
11509.
In addition to the original description? the genitalia have been illustrated by Betten (1934, pl. 22? fig. 8? and pl. 23? fig. I).
Nyctiophylax fraternus (Banks) ? 1905b9 p. 17. (Cyrnus)
Lectotgpe? f emale.-Plummer7s Island? Maryland, August 28. No. 11538.
A species of doubtful standing? placed as a synonym of vestitus by Milne (1935).
Nyctiophylax marginalis Banks, 1930b7 p. 231? fig. 15. Lectotype, male.-Put-in-Bay, Ohio? August 5, 1926? G. Townsend. No. 16325.




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19381 North American Caddis Flies 13
The male genitalia of this species are very distinctive, especially the sclerotized point on the mesa1 side of the clasper. They have been illustrated by Mosely under the name Cyrnellus ze~nii Mosely (1934? p. 142), which becomes a synonym of marginalis.
Nyctiophylax mcestus Banks, 1911, p. 359. Lectotype, mule.-Peachland? British Columbia, August 19, 1909, J. B. Wallis. No. 11536.
A synonym of vestitas Hagen.
*
Nyctiophylax vestitus ( Hagen) ? 1861? p. 293. (Polycen- tropus)
Lectotype, female.-Washington, D. C.? Osten Sacken. No. 11036.
The color and venation of this specimen associates it indubitably with the concept of this species as used by Banks? Betten and others.
Phylocentropus Iucidus (Hagen), 1861, p. 294. (Polgcen- tropm)
Lectotype, male.-Trenton Falls, New York, 1858, Osten Sacken. No. 11037.
The genitalia of this species also have been figured by Betten (1934, pl. 23, figs. 9-14).
Phylocentropus placidus (Banks), 1905b, p. 15. (Holocen- trop%s)
Lectotype, mule.-Washington, D. C.? August 25. No. 11540.
The essential features of the genitalia of this species have been illustrated by Betten (1934? pl. 23, figs. 2-18]. The species carolinus Carpenter has been placed as a synoHym erroneously by Milne.
Polycentropus albipunctus (Banks) ? 1930a, p. 131, figs. 6, 9. (Plectrocnemia)
Lectotype, mule.-Point Brevis, Cape Breton Island? Nova Scotia, July lo? 1928. No. 16324. LectoulLotype, female.- Same data.
The genitalia of the male of this species are shown in fig. 17.
Polycentropus arizonensis Banks, 1905b, p. 16. Lectotype, male.-Huachua Mts.? Arizona, June 21. No.



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14 Psyche [March
11546. Lectoallotype, female.-Same but July 20, 1903. The genitalia of this distinctive species are shown in fig. 15.
Polycentropus cinereus Hagen.-see Plectrocnemia cine rea (Hagen)
Polycentropus lucidus Hagen.-see Phylocentropus lucidus ( Hagen 1
Polycentropus remotus Banks, 1911, p. 359. Lectotype, male.-Peachland, British Columbia, August 23, 1909, J. B. Wallis. No. 11549.
The male genitalia are shown in fig. 16. Polycentropus vestitus Hagen.-see N~ctiophylax vestitus (Hagen)
Psychomyia diversa (Banks), 1914, p. 253, fig. 64. Lectotype, male.-Black Mountain, North Carolina, May, along north fork Swannanoa River. No. 11533. This is close but distinct from griselda (Betten). Psychomyia flavida Hagen.-see Psychomyiella jlavida (Hagen)
Psychomyia pulchella Banks.-see Psychomyiella pulchella (Banks)
Psychomyiella flavida (Hagen) , 1861, p. 294. (Psychomyia) Lectotype? female.-St. Lawrence River? Canada, 1859? Osten Sacken. No. 11055.
The structure of the male genitalia, fig. 19, indicates that this species belongs to Psychomyiella Martynov, and that Qwissa Milne? with jlavida as its genotype, is a synonym of it. The female genitalia are distinctive, allowing certain determination of this type. The species is widely distributed. -
PsychomyieIIa pulche~la (Banks), 1899, p. 217. Leetotype, male.-Colorado, accession no. 2022. No. 11534. A synonym of flavida (Hagen).
Family HYDROPSYCHIDB
Arctopsyche grandis (Banks), 1900a, p. 258. (Hydropsyche) Lectotype, male.-South West Colorado, July 20, 1899. No. 11514.




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19381 North American Caddis Flies 15
The mottled wings and male genitalia, fig. 36, will serve to distinguish this species from other nearctic members of the genus.
Cheumatopsyche analis (Banks), 1903b, p. 243. (Hgdro-
pwche)
Lectotype, male.-Riverton, New Jersey, July 16. No.
11532.
The elongate apical segment of the clasper, the V-shaped ridge on the dorsum of the tenth tergite, and the inconspic- uous apical lobes on the tenth tergite, fig. 20, distinguish the species from others in the genus. To date I have seen no specimens of this species except the type. The much used name "Hydropsychodes analis" has been applied to at least six different species in the past, and records under this name can not be accepted. Cheumatopsyche gracilis (Banks), 1899, p. 216. (Hydro- ~Whe)
Lectotype, male.-Colorado, accession no. 2022. No. 11497. The elongate apical segment of the clasper combined with the elongate-trapezoidal apical lobes of the tenth tergite will serve to distinguish this species, fig. 23. Cheumatopsyche minuscula (Banks), 1907a, p. 130, pl. 8, fig. 5. (Hydj-opsyche)
Lectotype, male.-Plummers Island, Maryland, August 29. No. 11530, Lectoallotype, female.-Same data. The pointed and approximate apical lobes of the tenth tergite? fig. 21, set this species off at once from sordida (Hagen), with which it was incorrectly synonymized by Milne.
Cheumatopsyche sordida (Hagen), 1860, p. 285. (Hydro-
~s~che)
Lectotype, male.-St. Lawrence River? Canada, 1859, Sacken. No. 11015.
This black species is distinguished by the widely separated apical lobes of the tenth tergite; these lobes have a dorsal, truncate apex, fig. 24.
Cheumatopsyche speciosa
(Banks)? 1904d, p. 214, PI. 2,
fig. 6. (Hydropsyche)




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16 Psych9 [March
Lectotype, male.-Plummers Island, Maryland, August 28. No. 11502. Lectoallotype, f emale.-Same data. In addition to the three large yellow spots of the front wings, the genitalia are distinctive of the species (Betten, 1934, pi. 20, fig. 14).
Diplectrona modesta Banks, 1908b, p. 266, pi. 19, fig. 13. Lectotype, male.-Riverside, Massachusetts, June 4, C. W. Johnson. No. 11523. Lectoallotype, female.-Same data. The details of the male genitalia have been illustrated by Betten (1934, pi. 17, figs. 3-7).
Hydropsyche analis Banks.-see Cheumatopsyche analis (Banks)
Hydropsyche bifida Banks, 1905b, p. 15, fig. 14. Lectotype, male.-Colorado, accession no. 2175. No. 11503. Lectoallotype, female.-Colorado, accession no. 2135, Fort Collins, June.
The male genitalia of this species, fig. 32, are distinguished by the short, widely separated apical processes of the tenth tergite and the small spur at the end of the lateral processes of the sedeagus. The species is widely distributed across the continent.
Hydropsyche californica Banks, 1898, p. 217. Lectotype, male.-Tahoma, California, August 28, 1897. No. 11304.
The somewhat moniliform apex of the sedeagus distin- guishes this species from others in the scalaris group, fig. 26. Hydropsyche chlorotica Hagen, 1861, p. 290. Lectotype, male.-St. Lawrence River, Canada, 1859, Sacken. No. 11016.
This specimen is identical in genitalia with the lectotype of morosa Hagen recently erected by Banks (1938). Diag-
nostic features include the flattened spur at the end of the lateral arm of the sedeagus, and the relatively simple apex of the sedeagus, fig. 33.
Hydropsyche cockerelli Banks, 1905b, p. 14. Lectotype, male.-Pecos, New Mexico, June 26, M. Grab- ham. No. 11506. Lectoallotype, f emale.-Same, August 14, at light, Cockerell.




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19381 North American Caddis Flies 17
The long, curved spur at the end of the lateral process of the sedeagus, combined with the short, upright and notched apical processes of the tenth tergite, set off this species from its close relatives, fig. 34.
Hydropsyche depravata Hagen, 1861, p. 290. Allotype, male.-Georgia, 1860, Gerhard.
This specimen agrees perfectly in color, habitus and labels with the female type of the species. There is no doubt that it is the same species. The genitalia are quite distinct, fig. 31, and in a large number of points suggest that this species and its close allies form the most primitive stock in the genus. Hydropsyche divisa Banks.-see Smicridea divisa (Banks) Hydropsyche gracilis Banks.-see Cheumatopsyche gracilis (Banks)
Hydropsyche grandis Banks.-see Arctopsyche grandis (Banks)
Hydropsyche hageni Banks, 1905b, p. 14, figs. 6,10,12. Lectotype, male.-Travilah, Maryland, July. No. 11996. The greatly lengthened and flattened apico-lateral plates of the sedeagus set the species off from others of the scalaris group easily, fig. 22.
Hydropsyche incommoda Hagen, 1861, p. 290. Lectotype, male.-Georgia, Winthem. No. 11028. This species is a member of the scalaris group. The
structures at the apex of the sedeagus are shown in fig. 29. Hydropsyche kansensis Banks.-see Potom kansensis (Banks)
Hydropsyche minuscula Banks.-see Cheumatopsyche mi- nuwda (Banks)
Hydropsyche novamexicana Banks, 1904a, p. 110, fig. 12. Lectotype, male.-Roswell, New Mexico, August 22, Cockerell. No. 11505.
This species is a synonym of occidentalis Banks. Hydropsyche occidentalis Banks, 1900a, p. 258. Lectotype, male.-Pullman, Washington, August 7, 1898. No. 11500.
This species is closest to scalaris and venularis, but differs



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, 18 Psyche
[March
from both in details of the genitalia, fig. 27. Hydropsyche oslari Banks, 1905b, p. 13, fig. 2. Lectotype, male.-South West Colorado, July 23, 1899.
No. 11501. Lectoallotype, female.-Same, but July 17. The long, apical segment of the claspers and the short, approximate apical processes of the tenth tergite are diag- nostic for this species, fig. 35.
Hydropsyche partita Banks, 1914, p. 252, figs. 58, 59. Lectotype, male.-Switzers Camp, San Gabriel Mts., California, June. No. 11498.
This is a synonym of oslari.
Hydropsyche phalerata Hagen, 1861, p. 287. Allotype, male.-Great Falls, Virginia, July 27. The great similarity between the pinned lectotype and this allotype leaves no doubt of the correct association of the two. The upturned apex of the tenth tergite and the slightly enlarged apex of the sedeagus are diagnostic, fig. 25. Hydropsyche recurvata Banks, 1914, p. 253, fig. 73. (H. slossonse var.)
Lectotype, male.-Go Home Bay, Ontario, Split Rock, June 9, E. M. Walker. No. 11507.
This species is distinguished by the combination of a long, curved spur at the end of the lateral process of the sedeagus and the apex of the sedeagus being developed into a pair of large, lateral lobes. The species codona Betten (1934, pi. 18, figs. 10-12) is a synonym of recurvata.
Hydropsyche slossonae Banks, 1905b, p. 14, figs. 4,7. Lectotype, male.-Franconia, New Hampshire. No. 11495. Lectoallotype, female.-Same data. In general appearance this species resembles other mem- bers of the "alternans" group, but is distinguished from all others by the male genitalia, fig. 30. Diagnostic characters include: simple, spine-like spur on end of lateral process of aedeagus, three large pockets of spines within apex of sedeagus, and long, excavated processes of tenth tergite which form an apical horseshoe.
Hydropsyche sordida Hagen.-see Cheumatopsyche sordida ( Hagen)




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19381 North American Caddis Flies 19
Hydropsyche speciosa Banks.-see Cheumatopsyche speciosa (Banks)
Hydropsyche venularis Banks, 1914, p. 252, fig. 62. Lectotype, male.-Washington, D. C., June 22. No. 11508. A member of the scalaris group, characterized by the greatly enlarged head at the apex of the eedeagus, which is half again as deep as the stalk, fig. 28. Macronema carolina Banks, 1909, p. 342.
Lectotype, male.-Southern Pines, North Carolina, June, 1924, A. H. Manee. No. 11529.
This species, although widely distributed, always occurs south of the range of zebratum Hagen.
Macronema flavum Hagen.-see Potomyia flava ( Hagen) Macronema zebratum Hagen, 1861, p. 285.
Lectotype, male.-St. Lawrence River, Canada, 1859, 0. Sacken. No. 11027. Lectoallotype, female.-Same data. This northern species is larger than carolina Banks and has the basal antenna1 segment more bulbous. Potomyia flava (Hagen), 1861, p. 285. (Macronema) Lectotype, male.-St. Louis, Missouri, 1859, Sacken. No. 11026.
This species has been illustrated by Betten (1934, pi. 21, figs. 1-3).
Potomyia kansensis (Banks), 1905b, p. 15. (Hydropsyche)
Lectotype, female.-Douglas County, Kansas, July, elec- tric light. No. 11499.
This represents the female of flava (Hagen), and is a synonym of this name.
Smicridea divisa (Banks), 1903a, p. 244, pi. 4, fig. 12. (Hydropsyche)
Lectotype, male.-Salt River, Arizona, April, Oslar. No. 11528. Lectoallotype, female.-Same data, but April 10. This species is a synonym of McLach1an7s fasciatella. The male genitalia show more relationship to Potomyia than to any other genus in the family.
Family ODONTOCERIDB
Heteroplectron dissimilis Banks, 1897, p. 30.



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20 Psyche [March
Lectotype, female.-Sea Cliff, Long Island, New York, June. No. 11718.
A synonym of indecisum (Walker).
Heteroplectron rufa (Hagen) , 1861, p. 276. (Molanna)
Lectotype, male.-Trenton Falls, New York, 1858, Osten Sacken. No. 10956. Lectoallotype, f emale.-Same data. This species also is a synonym of indecisum (Wlk.) Molanna rufa Hagen.-see Heteroplectron rufa (Hagen) Nerophilus calif ornicus (Hagen) , 1861, p. 272. (Silo) Lectotype, f emale.-Calif ornia. No. 10994. Allotype, male.-Same data.
The allotype may have been of the original type series. It displays the same distinctive color pattern as the female. Silo californicus Hagen.-see Nerophilus californicus ( Hagen)
Family MOLANNIDB
Molanna cinerea Hagen, 1861, p. 276.
Lectotype, female.-St. Lawrence River, Canada, 1859, 0. Sacken. No. 10957.
Only fragments of head and legs, and a fairly complete set of wings are left.
Molanna flavicornis Banks, 1914, p. 261, fig. 46. Lectotype, male.-Husavick, Manitoba, July 2, 1916, J. B. Wallis. No. 11590. Lectoallotype, f emale.-Winnepeg, Manitoba, May 31, 1911, J. B. Wallis.
The diagnostic features separating this species from others in the genus have been given by Betten (1934). Family LEPTOCERIDB
Athripsodes albostictus (Hagen), 1861, p. 276. (Leptocerus) Lectotype, female.-America, September, Winthem. No. 10963.
At the present time no satisfactory characters have been found to separate the females of this group of species. For the present, therefore, it will be necessary to consider this species of doubtful identity.
Athripsodes dilutus (Hagen), 1861, p. 277. (Leptocerus)




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19381 North American Caddis Flies 21
Lectotgpe, male.-Chicago, Illinois, Osten Sacken. No.
10965.
This species is a very close relative of annalicornis (Stephens), but may be readily separated from it on the basis of the sedeagus having two internal, large spines, and the clasper having a pedunculate sclerotized process, fig. 42. Dilutus has been incorrectly synonymized with annulicornis; it is really distinct and seems to be the only name available for the species.
Athripsodes flavus (Banks), 1904d, p. 212, pi. 2, fig. 4. (Leptocerus)
Lectotype, male.-Falls Church, Virginia. No. 11572. Lectoallotype, female.-Same data.
The angulate basal process of the claspers illustrated in the original description serves to differentiate this species from its only close ally, ancylus (Vorhies) . Athripsodes futilis (Banks), 1914, p. 264, figs. 44, 49. (Leptocerus)
Lectotype, male.-Go Home Bay, Ontario, July 11, E. M. Walker. No. 11574. Lectoallotype, female.-Same data, but August 12.
The male genitalia of this type agree with those of a male from Europe determined by Hagen as annulicornis (Stephens) and answer in detail the illustrations of Mc- Lachlan. There seems no doubt but that futilis must be considered a synonym of annulicornis. The chief diagnostic characters are in the sedeagus, which has only a single, large, internal spine, and the clasper, which has a short, digitate, sclerotized process, fig. 41.
Athripsodes lugens (Hagen), 1861, p. 276. (Leptocerzis)
Lectotype, male.-St. Lawrence River, Canada, 1859, 0. Sacken. No. 10966. Lectoallotype, f emale.-Same data. This species, also, is a synonym of annulicornis. Athripsodes maculatus (Banks), 1898, p. 214. (Leptocerus) Lectotype, female.-Washington, D. C. No. 11576. The type matches that of transversus (Hagen) so per- fectly that there is no doubt that the two are the same species. Both type series were collected at Washington.



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22 Psyche [March
Athripsodes transversus (Hagen), 1861, p. 279. (Lepto- c m )
Lectotype, female.-Washington, D. C., O'sten Sacken. No. 10967. Allotype, male.-Washington, D. C., June 22. A large series of this species in the M. C. Z. shows that it is the common one in the vicinity of the type locality, and that the designated allotype is correctly associated with Hagen's type. The male genitalia of the allotype are shown in fig. 40.
Athripsodes variegatus (Hagen), 1861, p. 278. (Leptocerus) Lectotype, male.-Chicago, Illinois, Osten Sacken. No. 10964.
The genitalia of this species have been illustrated by Betten under the name aspinosus Betten (1934, pi. 31, figs. 5-10). Both variegatus and aspinosus are synonyms of resurgens (Walker), according to the identification of Banks and Milne.
Leptocella candida (Hagen), 1861, p. 280. (Setodes)
Lectotype, male.-Florida, May, 1858, Sacken. No. 10972.
No attempt is made here to determine the status of any species in this genus.
Leptocella coloradensis Banks, 1899, p. 215. Lectotype, male.-Colorado, accession no. 2059. No. 11582. Lectoallotgpe, female.-Colorado, accession no. 2022. Leptocella minuta Banks, 1900a, p. 257.
Lectotgpe, f emale.-Pullman, Washington, August 9, 1898. No. 11581. Allotype, male.-Same data, but August 19.
The allotype bears no type label but is undoubtedly of the type lot.
Lept~cella nivea (Hagen), 1861, p. 281. (Setodes) Lectotype, male.-St. Lawrence River, Canada, 1859, Sacken. No. 10969.
Leptocella pavida (Hagen), 1861, p. 282. (Setodes) Lectotype, female.-Washington, [D. C.] , 0. Sacken. No. 10970.
Leptocella stigmatica Banks, 1914, p. 262, fig. 48.



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19381 North American Caddie Flies 23
Lectotype, male.-Jemey Mts., New Mexico, July 20. No. 11583.
Leptocella texana Banks, 1905b, p. 19.
Lectotype, male.-Zavalla County, Nueces River, Texas, April 27, 1910, Hunter & Pratt. No. 11578. Leptocerus albostictus Hagen.-see Athripsodes albostictus ( Hagen)
Leptocerus americana (Banks), 1899, p. 215. (Setodes)
Lectotype, female.-Washington, D. C. No. 11567. Lectoallotype, male.-Same data.
This species was made the basis for a new genus Ymyia by Milne.
All characters except the genitalia, however, group it with the genotype of Leptocerus. Leptocerus dilutus Hagen.-see Athripsodes dilutus ( Hagen)
Leptocerus flavus Banks.-see Athripsodes flavus (Banks) Leptocerus futilis Banks.-see Athripsodes futilis (Banks) Leptocerus grandis
(Banks), 1907a, p. 128, pi. 8, fig. 4.
(Setodes)
Lectotype, male.-New Haven, Connecticut, June 23,1904, H. L. Viereck. No. 11564. Lectoallotype, f emale.-Falls Church, Virginia, June 21.
This species is a synonym of americana.
Leptocerus lugens Hagen.-see Athripsodes lugens (Hagen) Leptocerus maculatus Banks.-see Athripsodes maculatus (Banks)
Leptocerus transversus Hagen.-see Athripsodes trans- versus (Hagen)
Leptocerus variegatus Hagen.-see Athripsodes variegatus ( Hagen)
Mystacides canadensis Banks, 1924, p. 448, fig. 47. Lectotype, male.-Sherbrooke, Canada. No. 14852. This represents the banded form of longicornis (L.). Field observations indicate that the unhanded specimens are simply rubbed, since the bands are formed only by hairs.



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24 Psyche [March
Mystacides interjecta Banks, 1914, p. 262, figs. 2, 5. ( OEceth)
Lectotype, female.-Go Home Bay, Ontario, August 22, E. M. Walker. No. 11551.
This species is a synonym of longicornis (L.). (Ecetina disjuncta Banks.-see (Ecetis disjuncta (Banks) CEcetina flavida Banks.-see OEcetis flavida (Banks) (Ecetina floridana Banks.-see (Ecetis floridana (Banks) (Ecetina fumosa Banks.-see (Ecetis fumosa (Banks) (Ecetina interjecta Banks.-see Mystacides interjecta (Banks)
CEcetina parvula Banks.-see (Ecetis parvula (Banks) (Ecetina persimilis Banks.-see OEcetis persimilis (Banks) (Ecetis cinerascens (Hagen), 1861, p. 282. (Setodes) Lectotype, male.-Washington, [D. C.] No. 10971. This species is illustrated by Betten (1934, pi. 35, figs. 2-7) under the name resurgens (Walker).
(Ecetis disjuncta (Banks), 1920, p. 351, fig. 100. (CEcetina) Lectotype, male.-Arroyo Seco Canyon, San Gabriel Mts., California, June 17, 1913, F. Grinnell, Jr. No. 10915. The male genitalia resemble those of avara (Banks) very closely.
(Ecetis flaveolata (Hagen), 1861, p. 282. (Setodes)
Lectotype, female.-Washington, [D. C.,] Osten Sacken. No. 10978.
This is the same species as incompicua (Walker) as determined by Milne.
(Ecetis flavida (Banks), 1899, p. 216. (OEcetina) Lectotype, male.-Kissimmee, Florida. No. 11557. Lectoallotype, f emale.-Same data.
This is the same species as inconspicm (Walker) as determined by Milne.
(Ecetis floridana (Banks), 1899, p. 216. (OEcetina) Lectotype, male.-Biscayne Bay, Florida. No. 11555. This is the same species as inconspima (Walker) as determined by Milne.




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19381 North American Caddis Flies 25
(Ecetis fumosa (Banks), 1899, p. 216.
(CEcetina)
Lectotype, f emale.-Washington, D. C. No. 11556. This species is a synonym of cinerascens (Hagen). CEcetis immobilis (Hagen), 1861, p. 283. (Setodes)
Lectotype, male.-St. Lawrence River, Canada, 1859. No. 10977.
The peculiar claspers, illustrated by Betten (1934, pi. 34, figs. 4, 5) readily distinguish this species. (Ecetis micans (Hagen), 1861, p. 283.
(Setodes)
Lectotype, male.-Washington, [D. C.,] 0. Sacken. No. 10973. Lectoallotype, female.-Same data. This is a synonym of inconspicua (Walker) as determined by Milne.
CEcetis parva (Banks), 1907a, p. 130, pi. 9, figs. 24, 26. (Setodina)
Lectotype, male.-Kissimmee, Florida. No. 11562. The genitalia of this minute leptocerid are illustrated in fig. 39. They are extremely similar to those of avara except for the claspers, which shows that Setodina must be con- sidered at most a subgenus of CEcetis.
CEcetis parvula (Banks), 1899, p. 215.
(CEcetina)
Lectotype, female.-Washington, D. C.
No. 11554.
This is the same as inconspicm (Walker) as determined by Milne.
(Ecetis persirnilis (Banks), 1907a, p. 129. ((Ecetina)
Lectotype, male.-High Island, Maryland. No. 11552. Lectoallotype, f emale.-Same data, in coitu with lectotype. The distinctive male genitalia are illustrated by Betten (1934, pi. 34, fig. 12).
CEcetis sagitta (Hagen), 1861c, p. 284.
(Setodes)
Lectotype, female.-Florida, March, 1858, 0. Sacken. No. 10975.
This is the same as inconspicua (Walker) as determined by Milne.
Setodes americana Banks.-see Leptocerus americana (Banks)
Setodes candida Hagen.-see Leptocella candida (Hagen)



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26 Psyche [March
' Setodes cinerascens Hagen.-see CEcetis cinerascens (Hagen 1
Setodes flaveolata Hagen.-see CEcetis flaveolata (Hagen) Setodes grandis Banks.-see Leptocerus grandis (Banks) Setodes immobilis Hagen.-see CEcetis immobilis (Hagen) Setodes injusta Hagen.-see Trisenodes injzbsta (Hagen) Setodes micans Hagen.-see (Ecetis mieans (Hagen) Setodes nivea Hagen.-see Leptocella nivea (Hagen) Setodes pavida Hagen.-see Leptocella pavida (Hagen) Setodes sagitta Hagen.-see (Ecetis sagitta (Hagen) Setodina parva Banks.-see OEcetis parva (Banks) Triaenodes borealis Banks, 1900a, p. 257. Lectotype, female.-St. Anthony Park, Minnesota. No.
11586.
Until definite characters are discovered for the lucid sepa- ration of the females of this genus, it will be necessary to consider this species of unknown status. Sufficient color antigeny exists in this genus to prevent the matching of males and females of each species on conventional characters alone.
Triaenodes dentata Banks, 1914, p. 261, fig. 45. Leetotype, male.-Johnstown, New York, June 28. No.
11589.
The male genitalia, fig. 38, are distinct in having the tenth tergite divided into a pair of long filaments, and having the lateral arm of the clasper of only medium length. Triaenodes flavescens Banks, 1900a, p. 257. Lectotype, male.-New Brunswick, New Jersey, October. No. 11588.
The details of the male genitalia have been illustrated under the name ignita (Walker) by Betten (1934, pi. 39, figs. 1-3).
Triaenodes frontalis Banks, 1907a, p. 127, pi. 9, fig. 11. Allotype, male.-Ft. Collins, Colorado, June 26, accession no. 2154.
This specimen was probably a part of the type series but did not bear a type label. The male genitalia differ from



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19381 North American Caddis Flies 27
those of grisea Banks chiefly in the long and whip-like mesal process of the clasper, fig. 37A.
Triaenodes grisea Banks, 1899, p. 214.
Lectotype, female.-Colorado, accession no. 2184. No.
11585. Allotype, male.-Denver, Colorado, June 27. This specimen was probably a part of the type series but did not bear a type label. The genitalia resemble those of frontalis, but differ in the short mesal processes of the claspers, fig. 37.
Triaenodes injusta (Hagen), 1861, p. 283. . (Setodes) Lectotype, male.-St. Lawrence River, Canada, 1859, Osten Sacken. No. 10976.
The genitalia of this species have been illustrated by Betten (1934, pi. 39, figs. 4-6).
Family PHRYGANEIDAZ
Agrypnia colorata Hagen, 1873, p. 424.
Lectotype, male.-Saskatchewan, 1860, Kennicott. No. 10734.
The male genitalia were illustrated by Milne (1931, figs. 10-11) under the name bradorata Milne.
Agrypnia straminea Hagen, 1873, p. 425.
Lectotype, male.-Saskatchewan, 1860, Kennieott. No. 10735.
The genitalia have been illustrated by Betten (1934, pi. 42, figs. 10-12) under the name mata Banks, which is a synonym of straminea.
Neuronia angustipennis Hagen, 1873, p. 400. Lectotype, male.-House, C [ambridge, Massachusetts ?I, July 9, 1863. No. 10739.
Neuronia stygipes Hagen, 1873, p. 388.
Lectotype, male.-West Roxbury, Massachusetts, April 26, 1868. No. 10741.
The male genitalia have been illustrated by Betten (1934, pi. 42, figs. 8, 9.
Family LIMNEPHILIDB
Acronopsyche pilosa Banks.-see Neophylax pilosus (Banks)




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28 Psyche [March
Anabolia assimilis Banks.-see Limnephilus assimilis (Banks)
Anabolia curta Banks.-see Limnephilus curtus (Banks) Anabolia modesta Hagen.-see Limnephilus modestus ( Hagen)
Anabolia montana Banks.-see Limnephilus montanm (Banks)
Anabolia nigricula Banks.-see Limnephilus nigricula (Banks)
Anisogamus costalis (Banks), 1901, p. 286. (Asynarchus)
Lectotype, male.-Las Vegas Range, New Mexico, June 28. No. 11676.
The generic placement of this species seems to me some- what doubtful. The male genitalia, fig. 44, show this species to belong to a residue of forms whose relationships are obscure.
Anisogamus disjunctus Banks, 1914, p. 156, fig. 22. Lectotype, male.-Bon Accord, British Columbia, May 22. Russell. No. 11673. Lectoallotype, female.-Same data, but May 18.
The male genitalia, fig. 43, lead to the same remarks as applied to the preceding.
Anisogamus edwardsi Banks.-see Druskws edwardsi (Banks)
Apatania canadensis Banks.-see Glyphopsyche canadensis (Banks)
Apatania pallida Hagen.-see Apatelia pallida (Hagen) Apatania shoshone Banks.-see Apatelia shoshone (Banks) Apatelia incerta (Banks), 1897, p. 28. (Enoicycla) Lectotype, male.-Sea Cliff, New York. No. 11681. Lectoallotype, female.-Franconia, New Hampshire. The male genitalia are illustrated in fig. 47. Milne (1935) considered this the same as nigra (Wlk.) . There are, how-
ever, two different species of Apatelia with long, needle-like claspers, and it is very likely nigra applies to one and incerta to the other.




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19381 North American Caddis Flies 29
Apatelia pallida (Hagen) , 1861, p. 270. ( Apatania) Lectotype, male.-St. Lawrence River, Canada, 1859, 0. Sacken. No. 14715.
This is a synonym of stigmatella (Zett.) . Apatelia shoshone (Banks), 1924, p. 442, figs. 35, 42. ( Apatania)
Lectotype, male.-Yellowstone Park, Wyoming, H. S. Smith. No. 14850. Lectoallotype, female.-Same data. The genitalic characters mentioned in the original descrip- tion will separate this species from its closest ally, stigmatella (Zett.) .
Apolopsyche pallida Banks.-see Limnephilus pallidus (Banks)
Asynarchus centralis Banks, 1900a, p. 253. Lectotgpe, male.-Clear Cr., Colorado, September 10,1899, Oslar. No. 11670.
The male genitalia, fig. 45, are suggestive of some species of Limnephilus, but other characters, such as the unusually long maxillary palpi, are quite distinctive. For the present it seems better to regard the generic assignment as open to question.
Asynarchus costalis Banks.-see Anisogamus costalis (Banks)
Asynarchus pallidus Banks, 1903b, p. 242. Lectotype, female.-South Park, Colorado, August 23, 1899, Oslar. No. 11671.
The species represents the opposite sex of contrails Bks. and is a synonym of it.
Asynarchus tristis Banks.-see Dicosmcecus tristis (Banks) Chilostigma subborealis Banks.-see Glyphopsyche sub- boreale (Banks)
Clistoronia maculata (Banks), 1904a, p. 107, pi. 1, figs. 2, 4. (Dicosmoecus)
Lectotype, male.-Pecos, New Mexico, August 24, at light, Cockerell. No. 11653. Lectoallotype, female.-South Arizona, August, 1902, F. B. Snow.
This species is the same as formosus (Banks). The male genitalia, fig. 46, show that the genus Clistoronia Banks is



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30 Psyche [March
closely allied to the Limnephilus section of the family, rather than to the Stenophylax section.
Colpotaulius medialis Banks.-see Limnephilw medialis (Banks)
Colpotaulius minusculus Banks.-see Limnephilus minus- culus (Banks)
Colpotaulius tarsalis Banks.-see Limnephilus tarsalis (Banks)
Dicosmoecus atripes (Hagen), 1875, p. 600. (Platyphylax) Lectotype, male.-Colorado Mts., August 1873, Comporte. No. 10701.
The male genitalia are illustrated in fig. 49. The color of legs is variable (see D. gilvipes) .
Dicosmoecus gilvipes (Hagen), 1875, p. 601. (Stenophylax)
Lectotype, male.-Quesnel Lake, British Columbia, August 27, Crotch. No. 10716.
The genitalia of this type are identical with those of atripes (Hagen). The color of the legs, previously used to separate the two, was found to vary over a large series of specimens. Gilvipes, therefore, becomes a synonym of atripes. Dicosmoecus maculatus Banks.-see Clistoronia maculata (Banks)
Dicosmoecus tristis (Banks) , 1900a, p. 254. ( Asynarchus ) Lectotype, male.-South Park, Colorado, August 17, 1899, Oslar. No. 11634. Lectoallotype, female.-Same data, but August 20.
This is a synonym of unicolor (Banks). The male genitalia differ considerably from those of atripes, fig. 48. Drusinus calypso (Banks), 1911, p. 350, pi. 13, fig. 25. (Stenophylax)
Lectotype, male.-Catskills, New York, June. No. 11672. This species is identical with sparsus (Bks) . Drusinus edwardsi (Banks), 1920, p. 345. (Anisogamus) Lectotype, male.-Marin County, California, H. Edwards. No. 10881. Lectoallotype, female.-California. The genitalia, illustrated in the original description, and general structure place this species in Drusinus Betten.



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19381 North American Caddis Flies 31
Ecclisomyia maculosa Banks, 1907a, p. 123, pi. 9, fig. 18. Lectotype, female.-Boulder, Colorado, July 31, 1904, Oslar. No. 11680. Allotype, male.-Slate Cr., Summit County, Colorado, August 2, 8, 211 ft. elevation, S. C. Clagg. The male genitalia, fig. 50, are quite distinctive. Enoicycla incerta Banks.-see Apatelia incerta (Banks) Enoicycia lepida Hagen.-see St enop hylax lepidus (Hagen) Glyphopsyche bellus (Banks), 1903b, p. 241. (Glypho- taeliw )
A male in the M. C. Z. identical in external characters with the female type is here designated the allotype. The male genitalia, fig. 51, are quite distinctive. Allotype, male.-March 23, 1903.
Glyphopsyche bryanti Banks, 1904c, p. 141. Lectotype, male.-Wellington, British Columbia, Bryant. No. 11640. Lectoallotype, f emale.-Same data. This lectotype is identical with paratypes of intercisus (Wlk.) in the M. C. Z., which species in turn has been synonymized with irroraAus (Fabricius) , fig. 53. Glyphopsyche canadensis (Banks), 1924, p. 442, fig. 5. ( Apatania)
Lectotype, male.-Winnipeg, Manitoba, October 1, 1909, J. B. Wallis. No. 14851.
The genitalia, fig. 52, are similar in general conformation to those of other nearctic members of the genus. Glyphopsyche subboreale (Banks), 1924, p. 441, fig. 32. ( Chilostigma)
Lectotype, male.-Alaska. No. 14847. Lectoallotype, female.-Beaver Mts., Alaska, May 15,1917, A. B. Twitchell. The male genitalia are illustrated in fig. 54. Glyphopsyche taylori (Banks), 1904c, p. 140. (Halesus) Lectotype, f emale.-Wellington, British Columbia, Bryant. No. 11666.
The extremely striking appearance of this species, the genotype of Halesochila Banks, is the chief character sepa- rating it from Glyphopsyche. Since it agrees so well with this latter genus in structural characters, it seems advisable to reduce Halesochila to subgeneric rank.



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32 Psyche [March
Glyphotaelius bellus Banks.-see Glyphopsyche bellus (Banks)
Glyphotaelius hostilis Hagen, 1864, p. 814. Lectotype, male.-Saskatchewan, 1860, Kennicott. No. 10730. Lectoallotype, female.-Fort Resolution, Great Slave Lake, Canada, 1862, Kennicott.
This large, distinctive species with the incised wing apex needs no comment.
Goniotaulius coloradensis Banks.-see Limnephilus colora- densis (Banks)
Grammataulius praecox Hagen, 1873, p. 451. Lectotype, male.-Fort Resolution, Great Slave Lake, Canada, 1862, Kennicott. No. 10732.
The species has been synonymized with interrogationis Zett.
Halesus amicus Hagen.-see Platycentropus amicus (Hagen)
Halesus hostis Hagen.-see Platycentrepm hostis (Hagen) Halesus taylori Banks.-see Glyphopsyche taylori (Banks) Halesus mutatus Hagen.-see Limnephilus mutatus ( Hagen)
Hesperophylax alascensis (Banks), 1908b, p. 265, pi. 19, fig. 14. (Platwhylax)
Lectotype, male.-Sitka, Alaska. No. 11647. This species is very close to and may be the same as designatus (Walker) .
Hesperophylax consirnilis Banks, 1900a, p. 253. (Limne-
philus)
Lectotype, male.-South Park, Colorado, August 25, 1899, Oslar. No. 11612. Lectoallotype, f emale.-Same data but July 20.
Differentiated by the sharp tenth tergite, fig. 55, this species is close to magnus Banks.
Hesperophylax magnus Banks, 1918, p. 20, fig. 9. Lectotype, male.-Palmerlee, Cochise County, Arizona, Biedermann. No. 10075. Lectoallotype, female.-Stockton, Utah, July 22, Spaulding.




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19381 North American Caddis Flies 33
This species is quite distinct from the other nearctic species of the genus on the basis of the cereus (see original description). It has been considered a subspecies of designatus Wlk. erroneously by Milne (1935). Hesperophylax occidentalis Banks, 1908b, p. 265, pi. 19, fig. 16. (Platyphylax)
Lectotype, male.-Florrisant, Colorado, August 3, 1907, S. A. Rohwer. No. 11646. Lectoallotype, f emale.-Ft. Win- gate, New Mexico, August, Wingate.
A close ally but not a synonym of designatus (Wlk.) . Homophylax crotch! Banks, 1920, p. 345.
Lectotype, female.-Victoria, British Columbia, July, Crotch. No. 10877.
The striking wing pattern of yellow and dark brown will serve to identify this species.
Homophylax flavipennis Banks, 1900a, p. 255. Lectotype, male.-South Park, Colorado, August 23, 1899, Oslar. No. 11635. Lectoallotype, female.-Same data. The male genitalia show many points of similarity to those of Glyphopsyche.
Leptophylax gracilis Banks, 1900a, p. 252. Lectotype, female.-St. Anthony's Park, Minnesota. No. 11637.
The male was described by Betten (1934, pi. 44, figs. 2-5). This seems to be one genus of the Limnephilus group suffi- ciently distinct to be recognized. At least, it can be keyed out in both sexes without difficulty.
Limnephilus Leach
Inability to find satisfactory characters for segregating the females into the same groups as the males has led me to consider as only of subgeneric rank such groups as Anabolia, Anabolina, Colpotaulius, Goniotaulius and some others. Limnephilus abbreviates Banks, 1908b, p. 263, pi. 18, fig. 8. Lectotype, male.-Tabernash, Colorado, August, E. S. Tucker. No. 11625. Lectoallotype, female.-Same data. The male genitalia are quite distinctive, fig. 71. Limnephilus aequalis Banks, 1914, p. 150, figs. 14, 31. Lectotype, male.-Bon Accord, British Columbia, June 7, Russell. No. 11624.




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34 Psyche [March
This species is a synonym of harrimani Bks., fig. 82. Limnephilus americanus Banks, 1900a, p. 253. Lectotype, female.-Idaho, C. V. Piper. No. 11631. This is considered a synonym of sublunatus Prov., fig. 73. Limnephilus argenteus Banks, 1914, p. 152, fig. 13. Allotype, male.-Hardisty Island, Great Slave Lake, Canada, June 5, 1924, J. Russell.
The male genitalia, fig. 56, are very different from any described Nearctic species in the genus. Limnephilus assirnilis (Banks), 1908b, p. 262, pi. 19, figs. 9, 10. (Anabolia)
Lectotype, ma,le.-Prescott, Arizona. No. 11648. Lecto- allotype, female.-Same data, June 15, 1902, Olslar. The slender and branched lateral arms of the sedeagus, fig. 58, distinguish this species from its closest allies. Limnephilus bifidus Banks, 1908b, p. 263, figs. 11, 13-15. Lectotype, f emale.-Pullman, Washington, C. V. Piper. No. 11627.
The type series contains only females.
Limnephilus brevipennis (Banks), 1899, p. 209. (Steno- phylax)
Lectotype, male.-Colorado, Gillette. No. 11612. In this species the male genitalia, fig. 77, appear to have the claspers much reduced and the lobes of the tenth tergite again divided.
Limnephilus canadensis Banks, 1908b, p. 264, pi. 18, fig. 4. Lectotype, female.-Lava1 County, Canada, June 29. No. 11619.
Until more female characters are worked out for this genus, the identity of this species cannot be determined. Limnephilus clausus Banks, 1924, p. 440, fig. 56. Lectotype, male.-Long Lake, Colorado, Cockerell. No.
14844. Lectoallotype, f emale.-Same data. This species is closely related to kincaidi Bks., the chief difference being in the lateral arms at the base of the apical portion of the sedeagus, fig. 64.
Limnephilus cockerelli Banks, 1900b, p. 124. Lectot ype, f emale.-Top of Range between Sapello and



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19381 North American Caddis Flies 35'
Pecos Rivers, New Mexico, August 2, 1900, 11,000' elev., T. D. A. & W. P. Cockerell. No. 11613. Allotype, male.- Top Las Vegas Range, New Mexico.
The male genitalia show this species to be a close relative of rnoestus Bks. The chief differences are found in the pro- portions of the tenth tergite and cerci, and details of the asdeagus, fig. 84.
Limnephilus coloradensis (Banks), 1899, p. 208. (Gonio- taulius )
Lectotype, male.-Colorado. No. 11621.
The curious sedeagus, fig. 76, and flattened claspers group this species with kennicotti Banks, although the two are by no means the same.
Limnephilus concolor Banks, 1899, p. 207. Lectotype, male.-Tacoma, Washington, September 5, 1897. No. 11611. Lectoallotype, female.-Same data. The abdomen of the lectotype is missing, hence it is irn- possible at present to give the diagnostic characters for the species.
Limnephilus consimilis Banks.-see Hesperophylax con- similis (Banks)
Limnephilus curtus Banks, 1920, p. 345, fig. 57. (Anabolia)
Lectotype, male.-Massachusetts. No. 10874. Lectoallo- type, female.-White Mts., New Hampshire. This species has been synonymized with planifrons Kolenati.
Limnephilus diversus (Banks), 1903a, p. 244, pi. 4, fig. 5. ( Anabolina,)
Lectotype, male.-Prescott, Arizona, Oslar. No. 11649. Lectoallot ype, f male.-Same data.
The cerci, fig. 79, distinguish this species from its closest relative, productus Bks.
~imne~hiius elongatus Banks, 1920, p. 344, figs. 92, 94, 98, 99.
Lectotype, male.-Fort Resolution, Great Slave Lake, Canada, 1862, Kennicott. No. 10870.
This species is a close relative of sublunatus Prov., but is readily distinguished by characters of the male genitalia, fig. 72.




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36 Psyche [March
Limnephilus externus Hagen, 1861, p. 257. Lectotype, female.-North Red River, Canada, 1854, Uhler. No. 10727.
The wing pattern is distinctive for the species. The male
genitalia are illustrated in fig. 80.
Limnephilus flavastellus Banks, 1918, p. 20, pi. 1, figs. 16, 17. Lectotype, male.-Jones Collection. No. 10072. Lecto- allotype, female.-Wellington, British Columbia, Sep- tember 23, 1903.
This species is a synonym of externus Hagen. Limnephilus gravidus Hagen, 1861, p. 257. Lectotype, female.-California. No. 10723. Allotype, male.-Santa Cruz, California, May 20, 1905. The abdomen of the type is partly gone. A male agreeing well with the type and designated allotype has male geni- talia, fig. 75, differing from those of vastus Hagen chiefly in details of the tenth tergite.
Limnephilus hageni Banks, 1930b, p. 226, figs. 7-9. Lectotype, male.-Fort Resolution, Great Slave Lake, Canada, 1862, Kennicott, No. 16316. Lectoallotpe, female. -Same data.
Belongs to the sublunata group, differing from other mem- bers as follows, fig. 69: appendages of the genitalia abbre- viated, the preanal appendages with a small sclerotization near the apex on the dorsal margin, and the sclerotized part of the lateral arms of the aedeagus stockier. Limnephilus hyalinus Hagen, 1861, p. 258. Lectotype, male.-North Red River, Canada, 1858, Uhler. No. 10722.
The male genitalia have been illustrated by Betten under the designation "Limnephilus sp. 1" (1934, pi. 46, fig. 5 and pi. 47, fig. 7).
Limnephilus medialis (Banks), 1905b, p. 8, pi. 1, fig. 3. (Colpotaulius)
Lectotype, male.-Muskoka, Ontario, July, 1888, E. P. Van Duzee. No. 11656. Lectoallotype, female.-Ithaca, New York, electric light.
This species has been synonymized with consocia Wlk.



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19381 North American Caddis Flies 37
Limnephilus janus new name for
Limnephilus minusculus (Banks), 1924, p. 439, fig. 52. (Colpotaulius) .
Preoccupied by Limnephilus minus-
culus (Banks), 1906, p. 120.
(Stenophylax) .
Lectotype, male.-Tolland, Colorado, Dodds. No. 14842. Lectoallotype, f emale.-Same data.
The male genitalia, fig. 59, are distinctive. I do not believe that this Colorado species is the same as perpusillus Walker, as stated by Milne.
Limnephilus modestus (Hagen), 1861, p. 265. (Anabolia) Lectotype, male.-Labrador, 1858, Hugo Christoph. No. 10711.
The differences between the genitalia of this species and mutatus (Hagen) are found chiefly in the cerci, fig. 65. Limnephilus moestus Banks, 1908a, p. 62, pi. 2, figs. 4, 8, 10, 11.
Lectotype, male.-Grand Lake, Newfoundland, July 28, 1906, 0. Bryant. No. 11629. Lectoallotype, female.-Same data.
A close relative of harrimani Bks., this species is distin- guished by the shorter cerci and less ornamented lateral arms of the sedeagus, fig. 83.
Limnephilus montanus Banks, 1907a, p. 119. ( Anabolia) Lectotype, female.-Mt. Katahdin, Maine. No. 11661. This species has been synonymized with planifrons Kol. and cu~tus Banks. Better characters in the female sex of this genus will have to be discovered to determine this with certainty.
Limnephilus morrisoni Banks, 1920, p. 343, figs. 5,32 and 96. Lectotype, male.-Reno, Nevada, 1878, Morrison. No. 10873. Lectoallotype, f emale.-Sierra Nevada, 1876, Crotch.
The male genitalia are illustrated in fig. 81. The generic assignment of this species is unsettled. Limnephilus mutatus (Hagen), 1861, p. 267. (Hallesus)
Lectotype, female.-Labrador, Winthem. No. 10688. Allotype, male.-Labrador, 1865, Moeschler. The allotype matches the lectotype so perfectly in general



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38 Psyche [March
characteristics and color that there seems no doubt of the correctness of the association. The male genitalia, fig. 66, are distinctive.
Limnephilus nigriculus (Banks), 1908b, p. 262, pi. 19, fig. 11. (Anabolia)
Lectotype, male.-Clear Cr., Colorado, September 10, 1889, Oslar. No. 11664.
The male genitalia are illustrated in fig. 68. They form an intermediate step between modesta, fig. 65, and mutatus, fig. 66.
Limnephilus occidentalis Banks, 1908b, p. 264, pi. 18, figs. 1, 3.
Lectotype, male.-Tacoma, Washington, September 5, 1897. No. 11620. Lectoallotype, female.-Wellington, British Columbia, Bryant.
The male genitalia, fig. 60, have small cerci and broad claspers. The lateral arms of the sedeagus are a primitive type*
Limnephilus oslari Banks, 1907a, p. 121, pi. 9, fig. 19. Lectotype, f emale.-South Park, Colorado, August 25, 1899. No. 11632.
The color pattern shows this to be the same as externus Hagen.
Limnephilus pacificus Banks, 1899, p. 207. Lectotype, male.-Olympia, Washington, Kincaid. No. 11610. Lectoallotype, female.-Tacoma, Washington, Sep- tember 5,1897.
This species has been synonymized with sitkensis (Kolenati) .
Limnephilus nepus new name for
Limnephilus pacificus (Banks), 1900a, p. 254. (Steno- phylax) . Preoccupied by the preceding species. Lectotype, male.-Pullman, Washington. No. 11663. Lectoallotype, female.-Same data, May 4, 1898. The male genitalia, fig. 67, have a curious sedeagus which is a simple derivative of the modesta type (see figs. 65, 66). Limnephilus pallidus (Banks), 1924, p. 442, fig. 54. (Apolo- psyche)




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19381 North American Caddis Flies 39
Lectotype, male.-Winnipeg Lake, Canada, 1860, Kenni- cott. No. 14849.
This is a synonym of parvulus (Banks).
Limnephilus perjurus Hagen, 1861, p. 258. Lectotype, female.-Ins. Kenae. No. 10721. The type lacks the abdomen.
Limnephilus pudicus Hagen, 1861, p. 262. Lectotype, female.-Washington, D. C., Osten Sacken. No. 10720.
This is a synonym of submonilifer Wlk.
Limnephilus pulchellus Banks, 1908a, p. 63, pi. 2, figs. 5, 9. Lectotype, male.-Grand Lake, Newfoundland, July 28, 1906, 0. Bryant. No. 11626. Lectoallotype, female.-Same data.
This small species has male genitalia, fig. 78, which bear a marked resemblance to the vastus group.
Limnephilus roberti Banks, 1930b, p. 226, figs. 10-12. Lectotype, male.-Winnipeg Lake, 1860, Kennicott. No.
16318. Lectoallotype, female.-Same data. This species is a synonym of parvulus (Banks), fig. 61. The type of this latter species lacks the abdomen, but a topo- type specimen which probably belonged to the type series but did not bear a type label agrees perfectly with the lecto- type of roberti.
Limnephilus sansoni Banks, 1918, p. 19, fig. 8. Lectotype, male.-Banff, Alberta, Spray Lake, August 25, Sanson. No. 10089. Lectoallotype, female.-Banff, Au- gust 31.
Belongs to the sublunatus group, set off from other mem- bers of the group by the following characters, fig. 70 : Pre- anal appendages very wide, with an inner band of heavily sclerotized points; sclerotized portion of the side arms of the asdeagus subdivided at their apex into small projections, otherwise the sedeagus is like sublunatus. Limnephilus secludens (Banks), 1914, p. 152, figs. 17,27. Lectotype, male.-Saskatchewan, July. No. 11623. Close to tarsalis Banks, but separated on details of the male genitalia, fig. 63.




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40 Psyche [March
Limnephilus sordidus Hagen, 1861, p. 264. Lectotype, male.-North Red River, Canada, 1858, Uhler. No. 10710.
This species has been considered a synonym of bimacu- lat us Wlk.
Limnephilus spinatus Banks, 1914, p. 149, fig. 8, 9. Lectotype, male.-vineyard, Utah, August 22. No. 11617.
Lectoallotype, f emale.-Same data, but August 28. The slender and undivided lateral arms of the sedeagus distinguish this species, fig. 57.
Limnephilus tarsalis (Banks), 1920, p. 342, fig. 104. (Col-
potaulius)
Lectotype? male.-Ward, Colorado, Oslar. No. 10880. The male genitalia, fig. 62, resemble most closely those of secludens (Banks).
Limnephilus vastus Hagen, 1861, p. 257.
Lectot ype, male.-Ins. Kenae. No. 10724. The abdomen of the type is missing.
Neophylax fuscus Banks, 1903b, p. 242.
Lectotype, male.-Agricultural College, Michigan, Pettit. No. 11643. Lectoallotype, female.-Franconia, New Hamp- shire, Mrs. Slosson.
The male genitalia of this species are illustrated in fig. 86. The species considered and illustrated as this by Betten is quite different.
Neophylax occidentis Banks, 1924, p. 441, figs. 51, 58. Lectotype, male.-Reno, Nevada, 1878, Morrison. No. 14848.
The male genitalia of this western species are illustrated in fig. 85. The similarity of the fundamental pattern of genitalia with the eastern species, and having the distinctive wart behind the lateral ocellus, indicate that this species forms a primitive subgenus of Neophylax. Neophylax pilosus (Banks), 1930b, p. 228, fig. 13. (Acrono- P~Y the)
Lectotype, male.-Modoc Co., California, July 20, 1922, Lindsey. No. 16319. Lectoallotype, f emale.-Same data. This species, the genotype of Acronopsyche Banks, is a synonym of occidentis Banks.




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19381 North American Caddis Flies 41
Platycentropus amicus (Hagen), 1861, p. 265. (Halesus) Lectotype, f emale.-New Orleans. No. 10690. No definite placement of this species can be made until well associated males are found, or good structural char- acters for separating the females.
Platycentropus hostis (Hagen), 1861, p. 266. (Halesus) Lectotype, male.-North Red River, Canada, 1858, Uhler. No. 10689. Lectoallotype, f emale.-Data illegible. This species has recently been synonymized with indicans (Walker) by Milne (1936).
Platyphylax alascensis Banks.-see Hesperophylax alas- censis (Banks)
Platyphylax atripes Hagen.-see Dicosmoecus atripes ( Hagen)
Plat yph ylax occidentalis Banks.-see Hespero phylax occi- dentalis (Banks)
Pycnopsyche similis Banks.-see Stenophylax similis (Banks)
Stenophylax brevipennis Banks.-see Limnephilus brevi- pennis (Banks)
Stenophylax calypso Banks.-see Drusinus calypso (Banks) Stenophylax gilvipes Hagen.-see Dicosmoecus gilvipes ( Hagen )
Stenophylax lepidus (Hagen) , 1861, p. 269. (Enoicy cla) Lectotype, male.-Pennsylvania. No. 10697. The male genitalia have been figured by Betten (1934, pi. 50, figs. 4-6). Lepida was sunk by Milne as a synonym of subfasciata (Say) but is distinct from that species. Stenophylax pacificus Banks.-see Limnephilus pacificus (Banks)
Stenophylax similis (Banks), 1907a, p. 122, pi. 9, fig. 22. (Pycnopsyche )
Lectotype, male.-Chatham, Michigan, August 23, 1900. No. 11659.
The male genitalia show this to be a synonym of guttifer Walker.




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42 Psyche [March
Family SERICOSTOMATIDB
Alepomyia bryanti Banks.-see Lepidostoma bryanti (Banks)
Arcadopsyche prominens Banks.-see Lepidostoma promi- nens (Banks)
Atomyia modesta Banks.-see Lepidostoma modesta (Banks)
Brachycentrus incanus Hagen, 1861, p. 272. Lectotype, female.-Washington, D. C., April, 1859, 0. Sacken. No. 10455.
This is probably numerosus (Say) or lateralis (Say). To
date, however, diagnostic characters have not been found for the females of this genus.
Brachycentrus occidentalis Banks, 1911, p. 355, pi. 13, fig. 32. Lectotype, male.-Bon Accord, British Columbia, May 14. No. 11685.
The long, separate cerci and slender, angled claspers, fig. 88, are diagnostic for this western species. Brachycentrus similis Banks, 1907a, p. 124, pi. 9, fig. 21. Lectotype, male.-Tabernash, Colorado, August, E. S. Tucker. No. 11684. Lectoallotype, female.-Boulder, Colorado, August 9, at light, T. D. A. Cockerell. This species is a synonym of americanus Bks. The short, fused cerci and bilobate claspers, fig. 87, are diagnostic for this widespread species.
Dasystoma rusticum Hagen.-see Micrasema rusticum ( Hagen)
Helicopsyche arizonensis Banks, 1907a, p. 125. Lectotype, female.-Nogales, Arizona, July, 1903, Oslar. No. 11694.
Helicopsyche californicus Banks, 1899, p. 210. Lectotype, male.-Colton, California. No. 11696. The genitalia are remarkably similar to those of borealis (Hagen).
Helicopsyche borealis (Hagen), 1861, p. 271. (Notidobia) Lectotype, male.-St. Lawrence River, Canada, 1859, 0. Sacken. No. 10939.




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19381 North American Caddis Flies 43
This widespread species has been illustrated by Betten (1934, pi. 66).
Lepidostoma Rarnbur
Many of the species here placed in Lepidostoma have been considered previously as belonging to genera separated from Lepidostoma on the basis of male characters such as vena- tion, folds in the wing, structure of antennae, etc. The male genitalia show that in many cases these definitions cut across phylogenetic lines, grouping together species which are at most distantly related and separating species which are really extremely closely related. The male genitalia show further that many of the most distinct of these groups are, at the most, an offshoot of a group of species placed in an- other genus. I feel, therefore, that many of these genera are only artificial segregates of species without any considera- tion to relationships. For this reason, I am defining the genus Lepidostoma very broadly. I agree with Mr. Banks that a thorough revision of the entire world fauna of this group will be necessary to correctly evaluate the genera. Lepidostoma bryanti (Banks), 1908a, p. 65, figs. 1, 2, 13. ( Alepomyia)
Lectotype, male.-Grand Lake, Newfoundland, July 28, 1906, 0. Bryant. No. 11709.
This species is a synonym of wisconsinensis Vorhies. Lepidostoma carolina (Banks), 1911, p. 356, pi. 13, fig. 28. (Notiopsyche)
Lectotype, male.-Southern Pines, North Carolina, April 28, A. H. Manee. No. 11704.
This species belongs to the togatum group as evidenced by the genitalia, fig. 92.
Lepidostoma costalis (Banks), 1914, p. 265, pi. 10, fig. 34. ( Olemira )
Lectotype, male.-Woodworth's Lake, Fulton Co., New York, August 19, 1909. No. 11701. Allotype, female.- Same data.
The allotype is evidently of the type series but did not bear a type label. The species has been illustrated by Betten (1934, pi. 64, figs. 1-5).




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44 Psyche [March
Lepidostoma grisea (Banks), 1911, p. 357, pi. 12, figs. 17, 19, 22. (Phanopsyche)
Lectotype, male.-Woodworth's Lake, Fulton Co., New York, August 22. No. 11693.
Characteristics of this species have been illustrated by Betten (1934, pi. 64, fig. 6-12).
It is a member of the
wiseonsinensis group.
Lepidostoma modesta (Banks), 1905a, p. 217. (Atomyza)
Lectotype, male.-Black Mts., North Carolina, May. No. 11702.
The elongate processes of the male tenth tergite, fig. 93, will identify this species.
Lepidostoma prominens Banks, 1930a, p. 129, figs. 1, 8, 10. (Arcadopsyche)
Lectotype, male.-Cape North, Cape Breton Island, Nova Scotia, August 7, 1928. No. 16321. Lectoallotype, female. -Same data.
The female bears the same label as the leetotype but did not have a type label. The tenth tergite of the male, fig. 91, is diagnostic.
Lepidostoma stigma Banks, 1907a, p. 125, pi. 8, fig. 10. Lectotype, female.-Boulder, Colorado, August 9, at light, T. D. A. Cockerell. No. 11692.
The placement of this species requires first identification of its corresponding male.
Lepidostoma togatum (Hagen), 1861, p. 273. (Mormonia)
Lectotype, f emale.-St. Lawrence River, Canada, 1859, Osten Sacken. No. 10942.
The traditional interpretation of this species is very likely correct, but until better diagnostic characters are found for the females of this genus, there is no guarantee of it. Lepidostoma vernalis (Banks), 1897, p. 29. (Mormonia)
Lectotype, male.-Sea Cliff, Long Island, New York, March 28. No. 11687. Lectoallotype, f emale.-Same data. The male genitalia, fig. 90, are characterized by the tooth- beset claspers, hooked processes of the tenth tergite and a pair of setal brushes on the ninth tergite. Micrasema charonis Banks, 1914, p. 266, figs. 3.47, 51.



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19381 North American Caddis Flies 45
Lectotype, male.-Black Mts., North Carolina, May. No. 11698.
This species differs from rustica (Hagen) in the curved apex of the claspers and more elliptic sedeagus, fig. 95. Micrasema falcatum Banks, 1914, p. 265, fig. 52. Lectotype, male.-Great Falls, Virginia, May 12. No. 11697.
This is a new synonym of rusticum (Hagen). Micrasema rusticum (Hagen), 1868, p. 272. (Dasystoma) Lectotype, male.-Saskatchewan, Canada, 1860, Kenni- cott. No. 10938. Lectoallotype, female.-Same data. The details of the genitalia are illustrated in fig Mormonia togatum Hagen.-see Lepidostoma
(Hagen )
Mormonia vernalis Banks.-see Lepidostoma (Banks)
Neothremma alicea Banks, 1930b, p. 229, figs. 4, 5. 96.
togatum
vernalis
Lectotype, male.-Colorado, G. S. Dodds. No. 16320. Lectoallotype, f emale.-Same data.
The male genitalia are illustrated in fig. 89. Notidobia americana Banks, 1900a, p. 256. Lectotype, male.-Falls Church, Virginia, June. No. 11714.
This species appears to be the same as grisea (Bks.), described from the female.
Notidobia assirnilis Banks, 1907a, p. 124, pi. 8, fig. 8. Lectotype, male.-San Diego, California, G. H. Fields. No. 11715.
This species has been considered the same as griseola McL. but is distinct from it on the basis of the claspers, fig. 94. Notidobia borealis Hagen.-see Helicopsyche borealis (Hagen)
Notidobia lobata Banks, 1911, p. 356, pi. 12, figs. 18, 20. (Schizopelex)
Lectotype, male.-Woodworth's Lake, Fulton Co., New York, June 22, 1910.
No. 11712.
This is a synonym of distincta (Ulmer).




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46 Psyche [March
Notidobia moesta Banks, 1914, p. 264, fig. 12. (Psiloneura)
Lectotype, f emale.-Cambridge, Massachusetts, Septem- ber. No. 11717.
This represents the female of the above, and is a synonym of it.
Notiopsyche carolina Banks.-see Lepidost oma carolina (Banks)
Olemira costalis Banks.-see Lepidostoma costalis (Banks) Phanopsyche grisea Banks.-see Lepidostoma grisea (Banks)
Psiloneura moesta Banks.-see Notidobia mmta (Banks) Schizopelex lobata Banks.-see Notidobia lobata (Banks) Banks,
1895
1897
1899
1900a
b
1901
1903a
b
1904a
b
c
d
1905a
b
BIBLIOGRAPHY
Nathan
New Neuropteroid Insects.
Transactions American Entomo-
logical Society, 22: 313-16.
New North American Neuropteroid Insects. Transactions American Entomological Society, 24: 21-31. Descriptions of New North American Neuropteroid Insects. Transactions American Entomological Society, 25 : 199-218. New Genera and Species of Nearctic Neuropteroid Insects. Transactions American Entomological Society, 26 : 239-59. Some Insects of the Hudsonian Zone in New Mexico. Psyche,
9: 123-26.
Some Insects of the Hudsonian Zone in New Mexico. Neurop- teroid Insects. Psyche, 9: 286-87.
Neuropteroid Insects from Arizona. Proceedings Entomo- logical Society Washington, 5: 237-45, 1 pi. Some New Neuropteroid Insects. Journal New York Entomo- logical Society, 11 : 236-43.
Neuropteroid Insects from New Mexico. Transactions Amer- ican Entomological Society, 30: 97-110, 1 pi. Two Species of Hydroptilidae. Entomological News, 15: 116-17.
Two New Species of Caddice-Flies. Proceedings Entomological Society Washington, 6 : 140-42.
A List of Neuropteroid Insects, Exclusive of Odonata, from the Vicinity of Washington, D. C. Proceedings Entomological Society Washington, 6 no. 4: 211-17, 1 pi. Trichoptera, p. 211-17.
Descriptions of New Species of Neuropteroid Insects from the Black Mountains, N. C. Bulletin American Museum Natural History, 21 : 215-18.
Descriptions of New Nearctic Neuropteroid Insects. Trans-
actions American Entomological Society, 32: 1-20, pis. 1-2.



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North American Caddis Flies 47
Descriptions of New Trichoptera. Proceedings Entomological Society Washington, 8 nos. 3, 4: 117-32, pis. 8-9. New Trichoptera and Psocidae. Journal New York Entomo- logical Society 15: 162-66. Trichoptera, p. 162-64, figs. 1-5. Some Trichoptera and Allied Insects from Newfoundland. Psyche, 15, no. 4: 61-67, pi. 2.
Neuropteroid Insects-Notes and Descriptions. Transactions American Entomological Society, 34: 255-67, pis. 17-19. Two New Caddis Flies. Entomological News, 20: 342. Descriptions of New Species of North American Neuropteroid Insects. Transactions American Entomological Society, 37 no. 4. Trichoptera, p. 350-60, pis. 12-13. American Trichoptera-Notes and Descriptions. Canadian En- tomologist, 46: 149-56, 201-5, 252-58, 261-68, pis. 9, 10, 15, 20. New Neuropteroid Insects. Bulletin Museum Comparative Zoology Harvard, 62. Trichoptera p. 19-22, pi. 1. New Neuropteroid Insects. Bulletin Museum Comparative Zoology Harvard, 64: 299-362, 7 pis. Trichoptera, p. 342-62. Descriptions of New Neuropteroid Insects. Bulletin Museum Comparative Zoology Harvard, 65. Trichoptera, p. 439-55, pi. 1, 3, 4.
Trichoptera from Cape Breton, Nova Scotia. Bulletin Brooklyn Entomological Society, 25: 127-32, 10 figs. New Neuropteroid Insects from the United States. Psyche, 37: 223-33, 15 figs.
Notes on some Hydropsychidae. Psyche, 38: 126-130, 10 figs. Betten, Cornelius
1934 The Caddis Flies or Trichoptera of New York State. New
York State Museum Bulletin, no. 292: 576 pp., 61 text figs., 67 pis.
Hagen, Herman A.
1860 Die Phryganiden Pictet's nach Typen Bearbeitet. Stettiner Entomologische Zeitung, 21 : 274-90.
1861
Synopsis of the Neuroptera of North America, with a list of the South American Species. Smithsonian Miscellaneous Collections. 347 p. Trichoptera, p. 249-98, 328-29. 1864 Phryganidarum Synopsis Synonymica. Verhandlung Zoolo- gische Botanische Gesellschaft Wein, 14: 799-890.
1868 Monographie der Gattung Dasystoma Rambur. Stettiner En- tomologische Zeitung, 29 : 267-73.
1873 Beitrage zur Kenntnis der Phryganiden. Verhandlung Zoolo- gische Botanische Gesellschaft Wien, 23 : 377-452. 1875
Report on the Pseudo-Neuroptera Collected by Lieut. W. L. Carpenter in 1873 in Colorado.
Report United States Geo-
logical Survey of the Territories for 1873, p. 571-606. Milne, Lorus J.
1934-36 Studies in North American Trichoptera. Cambridge, Mass.,
128 pp., 2 pis.
Morton, Kenneth J.
1905 North American Hydroptilidae. New York State Museum Bulletin 86: 63-75, fig. 15 and pis. 13-15.



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Psyche
[March
Mosely, Martin E.
1934 Some new exotic Trichoptera. Stylops, 3: 139-142, 13 figs. Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 7.
Fig. 8.
Fig. 9.
Fig. 10.
Fig. 11.
Fig. 12.
Fig. 13.
Fig. 14.
Fig. 15.
Fig. 16.
Fig. 17.
Fig. 18.
Fig. 19.
Fig. 20.
Fig. 21.
Fig. 22.
Fig. 23.
Fig. 24.
Agapetus wmlleatus Banks, male genitalia. Glossosoma parvulum Banks, male genitalia. Rhyacophila atrata Banks, male genitalia; A, tenth tergite; B, sedeagus.
Rhyacophila brunnea Banks, male genitalia; A, tenth ter- gite; B, sedeagus.
Rhyacophila coloradensis Banks, male genitalia; A, tenth tergite ; B, asdeagus.
Rhyacophila acropedes Banks, male genitalia. Rhyacophila hyalinata Banks, male genitalia; A, tenth ter- gite; B, aedeagus.
Rhyacophila grandis Banks, male genitalia; A, tenth tergite. Rhyacophila invaria Walker, male genitalia; A, sedeagus. Hydroptila albicornis Hagen, male genitalia ; A, sedeagus ; B, claspers.
Hydroptila maculata (Banks), male genitalia. Stactobia pictipes (Banks), male genitalia; A, sedeagus. Plectrocnemia aureola Banks, male genitalia; A, same, ventral view.
Metrichia nigrita (Banks), male genitalia; A, wings. Polycentropus arizonensis Banks, male genitalia; A, same, dorsal view.
Polycentropus remotus Banks, male genitalia; A, tenth tergite.
Polycentropus albipunctus (Banks), male genitalia. Cernotina pallida (Banks), male genitalia; A, same, dorsal view.
Psychomyiella flavida (Hagen), male genitalia. Cheumatopsyche analis (Banks), tenth tergite; A, same, caudal view.
Cheumatopsyche mimiscula (Banks), sedeagus ; A, tenth tergite, dorsal view; B, tenth tergite, lateral view. Hydropsyche hageni Banks, apex of sedeagus, lateral and ventral views.
Cheumatopsyche gracilis (Banks), tenth tergite, caudal view; A, clasper, caudal view.
Cheumatopsyche sordida (Hagen), sedeagus ; A, tenth ter- gite, dorsal view; B, tenth tergite, lateral view.



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Fig. 25.
Fig. 26.
Fig. 27.
Fig. 28.
Fig. 29.
Fig. 30.
Fig. 31.
Fig. 32.
Fig. 33.
Fig. 34.
Fig. 35.
Fig. 36.
Fig. 37.
Fig. 38.
Fig. 39.
Fig. 40.
Fig. 41.
Fig. 42.
Fig. 43.
Fig. 44.
Fig. 45.
Fig. 46.
Fig. 47.
North American Caddis Flies
Hydropsyche phalerata Hagen, male genitalia; A, apex of sedeagus, ventral view.
Hydropsyche californica Banks, apex of sedeagus, ventral and lateral views.
Hydropsyche occidentalis Banks, apex of asdeagus, ventral and lateral views.
Hydropsyche venularis Banks, aedeagus, lateral and ventral views.
Hydropsyche incommoda Hagen, apex of sedeagus, ventral and lateral views.
Hydropsyche slossonse Banks, tenth tergite, lateral view; A, same, dorsal view; B, asdeagus.
Hydropsyche depravata Hagen, male genitalia; A, apex of sedeagus, ventral view.
Hydropsyche bifida Banks, asdeagus; A, tenth tergite, dorsal view; B, asdeagus, dorsal view.
Hydropsyche morosa Hagen, sedeagus.
Hydropsyche cockerelli Banks, male genitalia. Hydropsyche oslari Banks, sedeagus; A, male genitalia. Arctopsyche grandis (Banks), male genitalia. Trissnodes grisea Banks, male genitalia; A, Trknodes frontalis, clasper, lateral aspect.
Trisenodes dentata Banks, male genitalia; A, clasper, ventral view.
(Ecetis parva (Banks), male genitalia.
Athripsodes transversus ( Hagen), male genitalia. Athripsodes annulicornis (Stephens), clasper, caudal view. Athripsodes dilutus (Hagen), male genitalia; A, clasper, caudal view.
Anisogamus disjunctus Banks, male genitalia; A, sedeagus. Anisogamus costalis (Banks), male genitalia; A, sedeagus. Asynarchus centralis Banks, male genitalia; A, zedeagus. Clistoronia formosa (Banks), male genitalia; A, asdeagus. Apatelia incerta (Banks), male genitalia. Fig. 48. Dicosmoecus unicolor (Banks), male genitalia. Fig. 49. Dicosmoecus atripes (Hagen), male genitalia. Fig. 50.
Ecclisomyia maculosa Banks, male genitalia; A, clasper. Fig. 51. Glyphopsyche bellus (Banks), male genitalia; A, sedeagus; B, male genitalia, dorsal view.
Fig. 52. Glyphopsyche canadensis (Banks), male genitalia; A, same, dorsal view; B, aedeagus.




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50 Psyche [March
Fig. 53.
Fig. 54.
Fig. 55.
Fig. 56.
Fig. 57.
Fig. 58.
Fig. 59.
Fig. 60.
Fig. 61.
Fig. 62.
Fig. 63.
Fig. 64.
Fig. 65.
Fig. 66.
Fig. 67.
Fig. 68.
Fig. 69.
Fig. 70.
Fig. 71.
Fig. 72.
Fig. 73.
Fig. 74.
Fig. 75.
Fig. 76.
Fig. 77.
Fig. 78.
Fig. 79.
Fig. 80.
Fig. 81.
Glyphopsyche irroratus (Fabricius) , male genitalia; A, sedeagus.
Glyphopsyche subborealis (Banks), male genitalia; A, sedeagus; B, same, dorsal view of apex.
Hes-perophylax consirnilis Banks, male genitalia. Limnephilus argenteus Banks, male genitalia; A, sedeagus. PLATE VII
Limnephilus spinatus Banks, male genitalia; A, sedeagus. Limnephilus assimilis (Banks), male genitalia; A, sedeagus. Limnephilus janus Ross, male genitalia; A, asdeagus. Limnephilus occidentalis Banks, male genitalia; A, sedeagus. Limnephilus parvulus (Banks), male genitalia; A, sedeagus. Limnephilus tarsalis (Banks), male genitalia; A, sedeagus. Limnephilus secludens (Banks), male genitalia; A, sedeagus. Limnephilus clausus Banks, lateral arm of sedeagus. Limnephilus modestus (Hagen), male genitalia; A, sedeagus. Limnephilus mutatus (Hagen), male genitalia ; A, sedeagus. Limnephilus nepus Ross, male genitalia; A, sedeagus. PLATE VIII
Limnephilus nigriculus (Banks), male genitalia; A, asdeagus.
Limnephilus hageni Banks, male genitalia; A, sedeagus. Limnephilus sansoni Banks, male genitalia; A, sedeagus. Limnephilus abbreviatus (Banks), male genitalia; A, sedeagus.
Limnephilus elongate Banks, male genitalia; A, sedeagus. Limnephilus sublunatus Provancher, male genitalia; A, sedeagus.
Limnephilus vastus Hagen, male genitalia; A, same, dorsal view; B, lateral arm of sedeagus.
Limnephilus gravidus Hagen, male genitalia ; A, same, dorsal view; B, lateral arm of sedeagus. Limnephilus coloradensis (Banks), male genitalia; A, B, sedeagus.
Limnephilus brevipennis (Banks), male genitalia; A, . sedeagus.
Limnephilus pulchellus Banks, male genitalia; A, same, dorsal view; B, sedeagus.
Limnephilus diversus (Banks), male genitalia; A, asdeagus. Limnephilus externus Hagen, male genitalia; A, sedeagus. hnephilus morrisoni Banks, male genitalia; A, sedeagus.



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No~th American Caddis Flies
Fig. 82.
Limnephilus harrimani Banks, male genitalia; A, zedeagus. Fig. 83.
Limnephilus mcestus Banks, male genitalia; A, zedeagus. Fig. 84.
Limnephik coclcerelli Banks, male genitalia; A, zedeagus. Fig. 85.
Neophylax occidentis Banks, male genitalia. Fig. 86.
Neophylax fwcus Banks, male genitalia, caudal view; same, ventral view.
Fig. 87.
Fig. 88.
Fig. 89.
Fig. 90.
Fig. 91.
Fig. 92.
Fig. 93.
Fig, 94.
Fig. 95.
Fig. 96.
Brachycentrus americanus Banks, male genitalia. Brachycentrus occidentalis Banks, male genitalia. Neothremma alicea Banks, male genitalia. Lepidostoma vernalis (Banks) , male genitalia ; A, same, dorsal view.
Lepidostoma prominens Banks, male genitalia. Lepidostoma ca~olina (Banks), male genitalia; A, tenth tergite, dorsal view.
Lepidostoma modesta (Banks), tenth tergite, lateral view. Notidobia a.ssimi1i.s Banks, clasper.
Mierasema charonis Banks, male genitalia; A, adeagus and tenth tergite, dorsal view.
Mierasema rusticum (Hagen), zedeagus and tenth tergite, dorsal view; A, tenth tergite, lateral view; B, clasper, lateral view; C, style of tenth tergite.



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52 Psyche [March
Psyche, 1938 VOL. 45, PLATE I.
Ross - North American Caddis Flies




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19381 North American Caddis Flies 53
Psyche, 1938
VOL. 45, PLATE 11.
Ross - North American Caddis Flies




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34
Psyche, 1938
Ross - North American Caddis Flies




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1
Ross - North American Caddis Flies




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56 Psyche [March
Psyche, 1938
VOL. 45, PLATE V.
Ross - North American Caddis Flies




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19381 North American Caddis Flies 57
Psyche, 1938
VOL. 45, PLATE VI.
Ross - North American Caddis Flies




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58 Psyche [March
Psyche, 1938
VOL. 45, PLATE VII.
MUTATUS .1
Ross - North American Caddis Flies




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19381 North American Caddis Flies 59
Psyche, 1938
VOL. 45, PLATE VIII.
Ross - North American Caddis Flies




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[March
Psyche, 1938
VOL. 45, PLATE IX.
Ross - North American Caddis Flies




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19381 North American Caddis Flies 61
Psyche, 1938
VOL. 45, PLATE X.
MODESTA
n
Ross - North American Caddis Flies




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