Lynn S. Kimsey and Richard M. Bohart.
A Synopsis of the Chrysidid Genera of Neotropical America (Chrysidoidea: Hymenoptera).
Psyche 87(1-2):75-91, 1980.

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A SYNOPSIS OF THE CHRYSIDID GENERA
OF NEOTROPICAL AMERICA
(CHRYSIDOIDEA, HYMENOPTERA)*
BY LYNN S. KIMSEY AND RICHARD M. BOHART
Department of Entomology, University of California, Davis 95616 The chrysidid fauna of Central and South America, including Mexico, differs considerably from that of North America. There are only a few species whose range extends into both regions. The genera Cleptes, Mesitiopterus, Microsega, Muesebeckidium, Pseudolopy- ga, Minymischa, Xerochrum, Microchridium, Hedychreides, Chry- sura, and Chrysurissa are not reported from South or Central America. A number of genera are not found north of Costa Rica, including Duckeia, Nesogyne and Gaullea. Trichrysis and Neo- chrysis have undergone extensive species radiation in the neotropics. The few species in these two genera occurring in North America are their northernmost representatives.
The faunas of Baja California, north central and northwestern Mexico seem to be essentially temperate in origin. The genera Parnopes, Spintharosoma and Ceratochrysis belong to this temper- ate fauna and are considered in this survey because they are found in Mexico. Several of the genera found only in the southwestern United States may be present in Baja California, Sonora and Chihuahua, Mexico, so specimens from this region should probably be run through the key to the North American genera (Bohart and Kimsey, 1980). The chrysidid fauna of the Caribbean islands, excluding Trinidad and Tobago, appears to be primarily temperate in origin, with only one endemic genus, Nesogyne.
The principal work on neotropical Chrysididae has been done by Juan Brkthes, August Brulle, Robert du Buysson, Peter Cameron, Alphonse Ducke, Felix Guerin-Meneville, Karl Krombein, Alexan- dro Mocsary and Maximilien Spinola. Lesser work has been done by R. Benoist, H. Bischoff, R. M. Bohart, E. T. Cresson, A. da Costa Lima, Andrea Dahlbom, Vicente Perez-D'Angello, Johann Fabri- cius, Claude Joseph (H. Janvier), L. S. Kimsey, Walter Linsenmaier and Flaminio Ruiz.
*Manuscript received by the editor August 26, 1980. Pu&e Ìö7:75.9 (1980). hup ttpsychu einclub orgtS7/87-075 html



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76 Psyche [vo~. 87
The two largest chrysidid genera in the neotropics are Trichrysis and Neochrysis, whose dominance roughly corresponds to Cerato- chrysis and Chrysis in the nearctic fauna. Detailed examination of Trichrysis in both hemispheres has revealed four distinct groups. Old World Trichrysis can be divided into two subgenera, based on whether the discoidal cell is complete (Trichrysis s.s.) or incomplete (Chrysidea). The New World Trichrysis can also be divided into two groups based on the same characteristic. However, New World Trichrysis are different from the Old World forms since they have a distinct and almost articulated telomere on the male genital capsule (fig. 24). This discrete telomere has not been found in old World Trichrysis (fig. 22).
We therefore propose the establishment of two new subgenera in the genus Trichrysis. An asterisk after a species name indicates that we have not seen the type.
Trichrysis Liechtenstein
Trichrysis (Caenochrysis) Kimsey and Bohart, new subgenus (derivation: Kaeno = new, Gr.)
Subgenerotype: Chrysis tridens Lepeletier Included species: amazonica Mocsary, azteca Mocsary, brasiliana Guerin, carinata Say*, divergens Cresson, gibbosa Mocsary, mathani Ducke*, mucronata Brullk*, nigropolita Bischoff, par- vula Fabricius, taschenbergi Mocsary, triangulifera Mocsary, and tridens Lepeletier*.
Discussion: Members of the subgenus can be recognized by the sometimes thin but completely sclerotized outer veins of the discoidal cell in the forewing (fig. 18), and the presence of a telomere on the paramere (fig. 24).
Trichrysis (Lorochrysis) Kimsey and Bohart, new subgenus (deriv- ation: Loron = strap, Gr.)
Subgenerotype: Chrysis doriae Gribodo
Included species: areolata Mocsary, armata Mocsary, boscii Buys- son, crotonis Buysson, doriae Gribodo,puberula Spinola, quadri- rimosa Mocsary, saussurei Mocsary.
Discussion: This subgenus is distinguished by a faint or incompletely sclerotized discoidal cell in the forewing, with at least the anterior vein faint or lacking (fig. 17) and the presence of a telomere on the paramere. In general the species are smaller than those of Caeno- chrysis.




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Kimsey & Bohart - Chrysidid Genera
Trichrysis (Trichrysis) Lichtenstein
Su bgenerotype: Sphex cyanea Linnaeus
Discussion: Trichrysis S.S. is an eastern hemisphere group, recog- nizable by the completely sclerotized discoidal cell and no telomere on the paramere (fig. 22).
Trichrysis (Chrysidea) Bischoff
Subgenerotype: Chrysis pumila Klug
Discussion: Chrysidea is also an eastern hemisphere group which lacks a telomere and has the forewing discoidal cell incomplete, since at least the anterior vein of the discoidal cell is lacking or faintly indicated.
Neochrysis Linsenmaier
Neochrysis is the largest genus in the neotropics. This genus was divided into four subgenera by Bohart (1966). We have further elaborated the characteristics of each subgenus and separated the neotropical species accordingly. The list of species under each subgenus may not be complete. Each list is based on species where we have seen the type or seen reliably determined material (type not seen indicated by an asterisk).
Neochrysis Linsenmaier, 1959. Generotype by original designa- tion, Chrysis punctatissima Spinola, 1840 (nec Villers, 1789) = Chry- sis carina Brullk., 1846.
T-I11 pit row obsolescent, pits, when discernible, no larger than other punctures (fig. 11); 4 distal teeth on T-111; frons without a cross carina or rarely a few traces of it, no strong carina leading back from brow to partly enclose midocellus; pronotum without a complete lateral carina; propodeum without a strong median projection just behind metanotum.
Partial list of included species: cameroni (Buysson), carina Brullk, deuteroleuca (Mocsary), inseriata (Mocsary), lecointei Ducke, mon- tezuma (Cameron), paraensis (Ducke).
Pleurochrysis Bohart, 1963, new name for Pleurocera Gukrin (nec Rafinesque, 18 18). Subgenerotype by monotypy, Chrysis viridis Guerin, 1842 (nec Oliver, 1790) = Chrysis bruchi Brkthes, 1902. T-I11 pit row usually well developed and forming a strong groove, pits larger than punctures, at least laterally (fig. 12); 4 distal teeth on T-I11 or rarely 6; frons with or without a cross carina (fig. 13),



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78 Psyche [vo~. 87
sometimes a strong carina leading back to partly enclose midocellus; pronotum without a complete lateral carina; propodeum without a projection behind metanotum.
Partial list of included species: acuta (Brkthes), alfkeni (Ducke), ameginoi (Brethes), ancilla (Buysson), bruchi (Brkthes), cavifrons (Brullk), charruana (Brkthes), chilicola (Mocsary), fasciifera (Bis- choff), lynchi (Brkthes), mendozana (Mocsary), missionera (Brkthes), post ica (Brullk), ypirangensis (Buysson). Exochrysis Bohart, 1963, Subgenerotype by original designation, Chrysis panamensis Cameron, 1888.
T-I11 pit row or groove well developed; 4 distal teeth on T-111; frons with a cross carina which branches back to enclose or partly enclose midocellus; pronotum with at most a partial lateral carina (fig. 5); propodeum with a strong median projection just behind metanotum (fig. 5). Partial list of included species: imperforata (Gribodo), leucophris (Mocsary), leucostigma (Mocsary), panamensis (Camer- on), spinigera (Spinola).
Ipsiura Linsenmaier, 1959. Subgenerotype by original designation, Chrysis marginalis Brullk.
T-I11 pit row or groove well developed; 2-6 distal teeth on T-111; frons sometimes with a cross carina but always with a carina leading back from brow to partly enclose midocellus; pronotum with a sharp and practically complete lateral carina or flange (fig. 4); propodeum without a projection behind metanotum (fig. 4). Partial list of included species: albibasalis (Mocsary), bisulcata (Ducke), brevispinosa (Ducke), ellampoides (Ducke), friesiana (Ducke), genbergi (Dahlbom)*, klugi Dahlbom, leucobasis (Mocsary), leuco- cheila (Mocsary), leucochiloides (Ducke), longiventris (Ducke), marginalis (Brullk), myops (Buysson), neolateralis Bohart, obidensis (Ducke), pilijirons (Cameron).
Argochrysis Kimsey and Bohart, new genus (derivation: Argos = white Gr.)
Generotype: Chrysis mesillae Cockerel1
Included species: mesillae (Cockerell), trochilus (Buysson) and a number of undescribed species.
Discussion: This genus is distinguished by F-I as long or longer than F-I1 in both sexes; tongue short, less than half as long as flagellum from basal fold to apex; mesopleuron with shallow transverse groove



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19801 Kimsey & Bohart - Chrysidid Genera 79 or no groove; propodeal teeth tapering and apically pointed (fig. 14); (fig. 7) leg joints, and apical rim of T-I11 often white or hyaline; paramere digitate and or lobate (fig. 19); cuspis broadly rounded, and digitus equal to or shorter than cuspis and apically rounded (fig. 19). Examination of the type species of Spintharosoma Zimmermann, Chrysis chrysonota Dahlbom, from the Ethiopian Region reveals that the American species previously placed in this genus are not congeneric.In S. chrysonota the tongue is long, the propodeal teeth clublike (fig. 15), paramere simple and rounded and cuspis slender and tapering (fig. 21), the male F-I is less than one third as long as F- 11, and the mesopleuron divided into two parts by a deep transverse groove. American species also differ from Spintharina (generotype Chrysis vagans Radovskowsky). Spintharina has two large teeth on the mesopleuron, lobate propodeal teeth (fig. 16), and male genitalia with simple unlobate paramere and long slender digitus and cuspis (fig. 20).
Gaullea Buysson
The genus Gaullea is an obscure and poorly known genus. Gaullea was originally described from a single female, collected in Chaco, Santiago del Estero, Argentina. We have seen an additional 2 females and 3 males from Santiago del Estero, Argentina, plus the type. This genus is characterized by the very short RS stub, lack of a discoidal cell in the forewing and denticulate rim of T-I11 (figs. 6, 10). Linsenmaier (1959) placed Gaullea near Euchroeus. However, after close examination of the wing venation we have placed Gaullea near Argochrysis, because a crease in the wing indicates that the marginal cell would be closed if the RS stub were extended. The closed marginal cell is characteristic of Chrysis and its relatives. The expanded and bladelike metanotum and denticulate rim of T-I11 suggests that Gaullea is closely related to Argochrysis. The male genitalia of Gaullea is similar to that of Chrysis, with a simple though somewhat apically lobate paramere, broad foliaceous digitus and slender setose cuspis (fig. 23).
BIOLOGY
Host records for the neotropical Chrysididae are incomplete at best and nonexistant for Nesogyne, Cleptidea, Amisega, Pseudo- spinolia and Gaullea. The Amiseginae are typically phasmatid egg parasites. The remaining genera are wasp and bee parasites (table 1).



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Psyche
[Vol. 87
Figs. 1-7, Lateral view of body. 1. Duckeia cyanea Costa Lima. 2. Adelphe mexicana Mocsary. 3. Cleptidea aurora (Smith). 4. Neochrysis (Ipsiura) marginalis (Brullk). 5. Neochrysis (Exochrysis) panamensis (Cameron). 6. Gaullea argentina Buysson. 7. Argochrysis mesillae (Cockerell).



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Kimsey & Bohart - Chrysidid Genera
8 Elampus
L./
9 Omalus
10 Gaullea
\ 11 Neochrysis (Neochrysis)
/
12 Neochrysis (Pleurochrysis)
14 Argochrysis &
15 Spintharosoma
16 Spintharina
17 Trichrysis (Lorochrysis) -
18 Trichrysis ( C a e n o c h r y ~ w
Figs. 8-12, Apical margin of tergum 111. 8. Elampus nitidus (Aaron). 9, Omalus butleri Bohart and Campos. 10. Gaullea argentina Buysson. 1 1. Neochrysis (Neochry- sis) carina (Brullk). 12. Neochrysis (Pleurochrysis) bruchi (Brethes). Fig. 13, Front view of male face of Neochrysis (Pleurochrysis) bruchi (Brethes). Figs. 14- 16, Anteroventral view of propodeal tooth. 14. Argochrysis mesillae (Cockerell). 15. Spintharosoma chrysonota (Dahlbom). 16. Spintharina vagans Radovskowsky. Figs. 17- 18, Forewing venation. 17. Trichrysis (Lorochrysis) doriae Gribodo. 18. Trichrysis (Caenochrysis) tridens (Lepeletier).




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84 Psyche [vo~. 87
Pseudospinolia-host not recorded in neotropics; Chile; 1 species. Argochrysis-Sphecidae: Ammophila (not recorded in neotropics); Mexico: Chihuahua and Baja California. 1 species. Gaullea-host not recorded; Argentina; 1 species. Trichrysis-Sphecidae: Trypoxylon, Eumenidae; widespread; 20 species.
Neochrysis-Sphecidae: Podium, Trypoxylon, Sceliphron; wide- spread; 32 species.
Subfamily Parnopinae
Parnopes-Sphecidae: Bembicini (prob. only on Microbembex in Mexico), northern Mexico, Sonora; 1 species. ACKNOWLEDGMENTS
We are very grateful to Ignacio and Raquel Cid del Prado for their work on translating the generic key into Spanish. Key to the Genera of Neotropical Chrysididae For brevity in the key the following abbreviations have been used: T for tergum, S for sternum and RS for radial sector. 1. Abdominal venter convex ............................. 2 Abdominal venter flat to concave ...................... 7 2. T-1-11 together dorsally subequal to or shorter than length of T-111-IV (fig. 3); pronotum with submedial transverse groove (fig. 3); upper frons with groove descending from midocellus (Cleptinae) ........................ Cleptidea Mocsary T-1-11 together dorsally much longer than length of T-111-IV (figs 1, 2); pronotum usually without transverse groove; upper frons without median groove (Amiseginae) ............ 3 3. Propodeum rounded posterolaterally (fig. l), pronotal lobes ..................
usually not reaching tegulae (fig. 1) 4
Propodeum toothed posterolaterally (fig. 2), pronotal lobes reachingtegulae ................................... 6 4. Pronotum medially depressed or grooved; S-I1 produced, keeled anteriorly; gena flanged; propodeum strongly produced, posterior face almost flat (fig. 1). ... Duckeia Costa L,ima Pronotum not medially depressed, S-I1 simple; gena not flanged; propodeum broadly rounded posteriorly. .............. 5



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19801 Kimsey & Bohart - Chrysidid Genera 85 .................
5. Scutum with notauli Amisega Cameron
.......... Scutum without notauli Mesitiop/erus Ashmead 6. Pronotum with transverse anterior groove, female with normal wings; propodeal tooth broad, subtriangular (fig. 2) ...... .................................. Adelphe Moscary Pronotum without transverse carina, female brachypterous, ...................
propodeal tooth slender and digitate
............................... Nesogyne Krombein 7. RS stub sclerotized for less than half marginal cell length (if vein is extended by its traces and wing crease to wing margin); outer veins of discoidal cell faint or barely indicated (fig. 6); tarsal ......................................
claws various 8
RS stub sclerotized for more than half marginal cell length (if vein is extended by its traces and wing crease to wing margin); outer veins of discoidal cell usually sclerotized (figs. 4, 5, 7); .....................................
claws simple 14
8. Last visible tergum spiculate along apical margin; claws simple .................................................. 9 Last visible tergum not spiculate along apical margin; claws ...............................
dentate (Elampinae) 10
9. Tongue exserted and reaching near base of abdomen, last visible tergum subapically depressed on either side of midline; females with three normally visible terga, males with four .......................
(Parnopinae) Parnopes Latreille
Tongue short, not exserted; last visible tergum not subapically depressed; both sexes with three normally visible terga ........................
(Chrysidinae) Gaullea Buysson
10. Hindtarsal claw long and slender, with a medial, nearly per- pendicular tooth; from with close punctation along inner eye margins (punctures less than one puncture diameter apart; .......................
scutellum not anteriorly carinate
...................... Hedychridium Abeille de Perrin Hindtarsal claw without a single, medial, nearly perpendicular ..................
tooth; other characteristics various 11
11. Medial vein essentially straight or weakly curved; hindtibia and usually midtibia with a definite pit on inner surface toward apex; female S-I11 usually with a nodular apicomedial projection .................................. Hedychrum Latreille Medial vein strongly arched; hindtibia and midtibia without a pit on inner surface; female S-111 without an apicomedial projec- ............................................. tion 12




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86 Psyche [vo~. 87
12. Medial cell with numerous dark microsetae; last tergum with punctate medially unnotched apical margin; face hollowed and crossridged; hindtarsal claws usually with two to five subsidiary teeth (rarely only one subsidiary) ............. Holopyga Dahlbom
..................................
Medial cell essentially asetose; last tergum apicomedially im- punctate and usually notched or compressed and snoutlike (figs. 8, 9); face hollowed without crossridging; hindtarsal claws various ..................................... 13 13. T-111 apicomedially notched (fig. 9), emarginate or simple not compressed and snoutlike; metanotum rounded, not conical or produced and bladelike; scutum impunctate or with punctures clustered along or between notauli, except chilensis (Mocsary) ........................... Omalus Panzer T-111 apicomedially compressed and snoutlike (fig. 8); metano- turn conical or produced and bladelike; scutum densely punctate ............................ Elampus Spinola 14. Radial sector ending before wing margin so that marginal cell is broadly open (fig. 4) .............................. 15 Radial sector and its faintly sclerotized remnant narrowly closing marginal cell or almost so (figs. 7, 17, 18) ..... 19 15. T-111 apical margin smooth, slightly prolonged medially, especi- ally in male, anterolateral angles of T-I not unusually sharp Pseudospinolia Linsenmaier
..........................
T-111 ending in four to six teeth; anterolateral angles of T-I ..................................
unusually sharp 16
16. Propodeum with strong median projection just behind metano- turn (fig. 5) ............ Neochrysis (Exochrysis) Bohart Propodeum without strong median projection just behind metanotum ...................................... 17 17. Pronotum with sharp and practically complete lateral carina or flange (fig. 4) T-111 pit row or groove well developed (as in fig. 12) ................... Neochrysis (Ipsiura) Linsenmaier Pronotum without a sharp or practically complete lateral carina or flange; T-111 pit row various ..................... 18 18. T-111 pit row obsolescent, pits, when discernible, no larger than other punctures (fig. 11); irons without cross carina, rarely with traces remaining and no carinae extending back toward ocelli ..................... Neochrysis S.S. Linsenmaier T-111 pit row well developed and forming a strong groove, at least



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Kimsey & Bohart - Chrysidid Genera
8 7
some pits larger than punctures (fig. 12); frons usually with cross carina and carinae extending back toward ocelli .... .................. Neochrysis (Pleurochrysis) Bo hart 19. Outer veins of discoidal cell faint (fig. 17) ...a,............ ............ Trichrysis (Lorochrysis) Kimsey and Bohart Outer veins of discoidal cell sclerotized (fig. 18) ......... 20 20. T-I11 not apicomedially notched, usually with three to six teeth ................................................ 21 T-111 apicomedially deeply notched, lacking subsidiary teeth, except on either side of notch, central Mexico north ... 22 21. T-111 ending in three or fewer teeth or angles .............. .......... Trichrysis (Caenochrysis) Kimsey and Bohart T-I11 ending in four or six teeth or angles ................. /
............................... Chrysis Linnaeus 22. Tibiae with pale basal markings and T-III rim whitish; or meta- notum conically produced (fig. 7) ...................... .................... Argochrysis Kimsey and Bohart Tibiae without pale markings and T-I11 rim not whitish; metano- turn not produced ............... Ceratochrysis Cooper Clave para les generos neotropicales de la familia Chrysididae 1. Region ventral del abdomen convexa .................. 2 Region ventral del abdomen plana o concava ........... 7 2. T-1-11 dorsalmente subiguales o mas cortos que el largo de T-III- IV (fig. 3); pronotum con hendidura submedia transversa (fig. 3); frente superior con una hendidura descendiendo del ocelo medio (Cleptinae) ............ Cleptidea Mocsary T-1-11 dorsalmente tan largos que el largo de T-111-IV (figs 1, 2); pronotum sin o con hendidura transversal; frente superior sin hendidura media (Amiseginae) ....................... 3 3. Propodeum redondeado posteriolateralmente (fig. l), lobulos .... pronotales usualmente sin alcanzar la tkgula (fig. 1) 4 Propodeum dentado posteriolateralmente (Fig. 2), lobulos pronotales alcanzando la tegula ...................... 6



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Psyche [vo~. 87
4. Pronotum en su parte media hundido o con hendidura; S-I1 forma una projecion angular anteriormente; gena con pro- jecion o borde; proprodeum evidentemente pronunciado ...... posteriormente, extreme posterior casi piano (fig. 1) Duckeia Costa Lima
................................
Pronotum sin depresion media; S-I1 simple; gena sin projecion alar; propodeum distintamente redondeado posteriormente .................................................. 5 5. Scutum con notauli .................. Amisega Cameron Scutum sin notauli ............... Mesitiopterus Ashmead 6. Pronotum con hendidura transversal; hembra con alas normales; diente propodeal ancho, subtriangular (fig. 2) ........... Adelphe Mocsary
...................................
Pronotum sin hendidura transversal; hembra con alas bra- ......... chypterous; diente propodeal alargado y angosto Nesogyne Krombein
.................................
7. RS fragment0 esclerotizado, menor que la mitad del largo de la celula marginal (si la vena es extendida por sus trazos y plieque del ala marginal); venas externas de la celula discoidal tenues o apenas visibles (fig. 6); ufias tarsales variables . 8 RS escleratizado mas de la mitad del largo de la celula marginal (si la vena es extendida por sus trazas y plieque de ala marginal); venas externas de la celula discoidal usualmente ...... escleratizado (figs 4, 5, 7); ufias tarsales simples 14
8. ultimo tergum visible, espiculado a lo largo del margin apical; ...............................
unas tarsales simples 9
ultimo tergum visible, no espiculado a lo largo del margin apical; ufias tarsales dentadas (Elampinae) ........... 10 9. Lengua extendida, alcanza cerca la base del abdomen; ultimo tergum visible, subapicalmente hundido en su parte media; hembras con tres normalmente visibles terga, machos con cuatro (Parnopinae) ................ Parnopes Latreille Lengua corta, no se extiende; ultimo tergo visible no subapical- mente hundido; ambos sexos con tres normalmente visibles terga (Chrysididinae) ................. Gaullea Buysson 10. Ufias de les tarsos posteriores largas y delgadas, con un diente medio perpendicular; frente con cerrada puntacion a lo largo del margin interno del ojo (puntacion menos que el diametro de una puntacion distantes); scutellum no anteriormente con saliente ............... Hedychridium Abeille de Perrin



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19801 Kimsey & Bohart - Chrysidid Genera 89 Ufias tarsales posteriores simples, diente medio perpendicular presente; otras caracteristicas varias ................. 1 1 11. Vena media esencialmente recta o dkbilmente curvada; tibia posterior y tibia media usualmente con un claro hoyo en su superficie interna, cerca de su apice; S-I11 de la hembra ..... usualmente con una proyeccion nodular medioapical ............................ Hedychrum Latreille Vena media fuertemente arqueada; tibia posterior y media hoyo sobre su superficie interna; S-111 de la hembra sin una proyeccion medioapical ............................ 12 12. Cklula media con numerosas microsetas obscuras; ultimo tergo con mediana puntacion, no alcanza el margen apical; cara concava y con estriaciones transversales; unas traseras usual- ..... mente con 2 a 5 dientes subsidiaries (raremente uno) ............................... Holopyga Dahlbom Cklula media esencialmente sin setas; ultimo tergo sin mediana puntacion apical, usualmente con muescas o comprimido similar a una proyeccion truncada (figs 8,9); cara concava sin ........ estriaciones transversales; ufias traseras varias 13
13. T-I11 de la hembra medio apical con muescas (fig. 9), emarginado o simple no comprimido o comprimido y sin proyeccion truncada; metanotumo redondeado, no conico o aplanado; scutum con puntacion cerrada a lo largo o entre notauli, except0 chilensis (Mocsary) ............ Omalus Panzer T-I11 de la hembra medio apical comprimido y con projection truncada (fig. 8); metanotum con proyeccion conica o aplana- da; scutum densamente punteado ...... Elampus Spinola 14. Sector radial termina antes del margen del ala, cklula marginal es ampliamente abierta (fig. 4) ........................ 15 Sector radial con cklula marginal angosta y debilmente esclera- ................ tizado o casi cerrada (figs. 7, 17, 18) 19
15. T-I11 margen apical liso, ligeramente prolongado, en su parte media, especialmente en machos; angulos anterolateral de T-I no extraordinariamente agudos ........................ .......................... Pseudospinolia Linsenmaier T-I11 termina en cuatro o seis dientes; angulos anterolaterales de T-I extraordinariamente agudo ..................... 16 16. Propodeum con fuerte proyeccion media justo atras del metano- turn (fig. 5) ............ Neochrysis (Exochrysis) Bohart



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90 Psyche [vo~. 87
Propodeum sin fuerte proyeccih media, justo atras del metano- turn ............................................. 17 17. Pronotum con borde agudo y practicamente completa carina lateral (fig. 4); T-I11 con hilera de hoyos bien desarrollados ................ formando una hendidura (como fig. 12) .................... Neochrysis (Ipsiura) Linsenmaier Pronotum sin borde agudo o completa carina; T-I11 hilera de ..................................... hoyosvarios 18 18. T-I11 hilera de hoyos casi ausentes, hoyos cuando son discern- ible~, no tan largos que otras puntaciones (fig. 11); frente sin carina cruzada, raramente con remanentes y sin extenciones .......................... de la carinae hacia 10s ocelos ........................ Neochrysis s. s. Linsenmaier T-111 con hilera de hayos bien desarollado y formando un fuerte surco, algunos hoyos tan grandes que una puntacion (fig. 12); frente usualmente con carina cruzada y carinae extendiendase hacia 10s ocelos ....... Neochrysis (Pleurochrysis) Bohart ....... 19. Venas externas de la cklula discoidal dkbiles (fig. 17) ............ Trichrysis (Lorochrysis) Kimsey and Bohart Venas externas de la celula discoidal esclerotizadas (fig. 18) . . ............................................. 20 20. T-I11 medio apical sin muescas, usualmente con tres, cuarto o ..
seis dientes .................................... : 21 T-111 medio apical profundamente con muescas, dientes subsi- darios ausentes, except0 sobre ambos lados de la muesca; central y norte de Mexico .......................... 22 ........ 21. T-I11 terminado en tres o algunos dientes o angulos .......... Trichrysis (Caenochrysis) Kimsey and Bohart ......... T-I11 terminado en cuatro o seis dientes o angulos ................................. Chrysis Linnaeus



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Kimsey & Bohart - Chrysidid Genera
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Les hymenopteres qui habitent les tiges de sonce aux environs de Quito. Ann. Ent. Soc. France 111:75-90.
BOHART, R. M.
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The genus Neochrysis in America north of Mexico. Bull. Brooklyn Ent. SOC. 63: 139- 144.
COSTA LIMA, A DA.
1936.
Sur un nouveau chryside: Duckeia Jubilaire E. L. Bouvier, pp. 173-175. COVILLE, R. E. AND P. L. COVILLE.
1980.
Nesting biology and behavior of Trypoxylon (Trypargilum) tenoctitlan in Costa Rica. Ann. Ent. Soc. Amer. 73:llO-119. JANVIER, H.
1933.
Etude biologique de quelques Hymenopteres du Chili. Ann. Sci. Nat. Zool., Paris ( 10)16(14):209-356.
KROMBEIN, K. V.
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A generic review of the Amiseginae, a group of phasmatid egg parasites and notes on the Adelphinae. Trans. Amer. Ent. Soc. 82:147-215. LINSENMAIER, W.
1959.
Revision der familie Chrysididae. Mitt. Schweiz. Ent. Ges. 32:l-232. PEREZ-D'ANGELLO, V.
1968.
Insectos asociados a 10s nidos de Hypodynerus lachesis (Lepeletier). Rev. Chilena Ent. 6: 13 1- 134.
RU'IZ, F.
1938. Observaciones biologicas de algunos insectos chilenos. Rev. Univ. 23:148-153.
1939. El Odynerus labiatus Hal. y su biologia. Rev. Chilena Hist. Nat. 42:97- 105.




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