Klaus Dumpert.
Camponotus (Karavaievia) texens sp. n. and C. (K.) gombaki sp. n. from Malaysia in comparison with the other Karavaievia species (Formicidae: Formicinae).
Psyche 92(4):557-573, 1985.

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CAMPONOTVS (KARA VAIEVIA) TEXENS SP. N. AND C. (K.) GOMBAKI SP. N. FROM MALAYSIA
IN COMPARISON WITH THE OTHER KARAVAIEVIA SPECIES (FORMICIDAE: FORMICINAE)*
Battelle-Institut, Abteilung Chemie und Okologie Am Romerhof 35, D-6000 Frankfurt am Main, Federal Republic of Germany
Two weaving species of Camponotus, collected in a rain forest in Malaysia in 1984 (Maschwitz, 1985), have turned out to be new species of the subgenus Karavaievia. These are described below as Camponotus (Karavaievia) texens and C. (K.) gombaki and are compared with the other species of the subgenus Karavaievia Emery (1925).
MATERIALS AND METHODS
The two colonies of Camponotus texens were found by U. Maschwitz: one of the Gombak Valley, about 27 km north of Kuala Lumpur, and the other one in the region of Kuantan, on the east coast of Malaysia. Of C. gombaki, only one colony was discovered in the Gombak Valley. Twenty workers of each of the colonies were used for the descriptions that follow. The description of the sexuals for C. texens was based on 5 males from the colony of Kuantan and 5 females from the Gombak Valley. Only 5 females, but no males, were available for C. gombaki.
One worker of Camponotus (Karavaievia) overbecki was made available to the author by Dr. Baroni Urbani from the Santschi Collection in Base1 (Switzerland). Three workers of this species were used for classifiction from the Fore1 Collection in Geneva (Switzer- land) along with three workers and two dealate females of Campo- notus (Karavaievia) exsectus from the Emery Collection in Genoa (Italy). However, specimens of C. (Karavaievia) dolichoderoides were not available from either the Fore1 or Emery Collections. *Final copy of manuscript received by the editor October 14, 1985. 557
Pu&e 92557.574 (1985). http:llpsyche cnlclub oreflW2-557 html



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330 rsycne [VOL 92
The examinations of C. texens, C. gombaki and C. overbecki were made with the aid of a binocular microscope; the measure- ments of the animals were made at a magnification of 65X. Scan- ning electron micrographs of C. texens and C. gombaki, presented as part of this article, were taken by a Hitachi S 500. The following abbreviations are used below (measurements as in Bolton, 1977):
TL: total length. The total outstretched length of the individual from the mandibular apex to the gastral apex. HL: head length. The straight-line distance between the anterior clypeal margin and the mid-point of the occipital margin in full-face view.
CI: cephalic index = (H W X loo)/ H L
SL: scapus length. The straight-line length of the antenna1 scape excluding the radicle.
SI: scapus index = (H W X loo)/ SL
PW: pronotal width. The maximum width of the pronoturn in dor- sal view.
OD: ocular diameter. The maximum width of the eyes. All measurements are expressed in millimeters. The critical reading of the manuscript by Dr. C. Baroni Urbani and Dr. P. van Voris is gratefully acknowledged. RESULTS AND DISCUSSION
Characteristics of the Subgenus Karavaievia Emery 1925 The description of the subgenus Karavaievia follows that of Emery (1925). This description is mainly a listing of characters which are partly superficial but which serve to distinguish this sub- genus from all the others (Emery, 1925). A thorough diagnosis of 'Karavaievia' would require a revision of the whole genus Campo- notus, which is urgently needed but which is not the intention of this paper. Thus, the subgenus Karavaievia is in a preliminary state like the classification of the genus Camponotus itself. According to Emery (1925) Karavaievia is closely related to the subgenera Myrmamblys, Myrmoteras, and Myrmoplatys. The most important difference between these subgenera and Karavaievia is the nearly complete absence of any dimorphism in the worker caste of the Karavaievia species. Specific for Myrmoplatys is the scapal



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insertion near the midlength of the frontal carinae, the depression of the anterior part of the head and the relatively short flagellar seg- ments. Myrmotarsus includes massive and large species with depressed tibiae. Workers of Myrmarnblys have heads that are longer than wide with parallel sides. The characteristics of Kara- vaievia are as follows:
1 Workers
Apart from mid& variability, all workers are monornorphic and about 5.5 mm long. Head trapezoidal, with rounded occipital corners. Head sides convex, occipital margin more or less concave, and anterior margin mostly straight. The eyes are situated behind the midlength of the sides of the head. Length of head subequal to its width (i.e. CI - 100). Clypeus narrow, convex, and without median carina. Its anterior margin almost straight, and in the mid- dle third slightly concave. Frontal carinae relatively short and sub- parallel to moderately divergent. Mandibles short, with lateral borders strongly curved and 5 teeth on each masticatory border. Antenna1 scapes projecting beyond the occipital margin by about one third of their length. Pedicel longer than the following flagellar segments; apical flagellar segments slightly thickened. Alitrunk with a deep impression (metanotum) between promesonotum and pro- podeum, and two raised stigmata at the deepest point of the impres- sion. Petiolar scale tapers to a transverse ridge. Females
Total length about 1 1 mm. Head about twice as wide as that of the workers, and with less convex sides. Anterior clypeal margin with a median semicircular excision. Apical flagellar segments not thickened .
Males
Anterior clypeal margin straight. Mandibles with only one strong, apical tooth. Total outstretched length about 6.5 mm. Camponotus texens sp. n.
Derivatio nominis: The species name is derived from the Latin word 'texere' = weave.
A. Holotype $: TL 5.4, HL 1.26, HW 1.29, CI 103, SL 1.51, SI 85, PW 0.77, OD 0.3 1.




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Frontal carinae extend to about midlength of head. Apart from a slight projection behind the scapal insertion, they are straight and slightly divergent (Fig. 1 a). Anterior clypeal margin almost straight, but in the middle third weakly concave, with slightly indented edges. Eyes are situated behind the midlength of the sides of the head; their maximum diameter is 0.31 mm, or about 0.24 HW. Head and alitrunk dark brown; gaster and petiole dirty yellow. Antennae and legs, including tibiae, dark brown, while apical flagel- lar segments, tarsi and front of head are dirty yellow like the gaster. Surface of head, alitrunk and gaster weakly shining. Accordingly, cuticular punctures and reticulated sculpture (SEM) are weak (Fig. la, c). Gaster with an imbricate sculpture (Fig. 3b), which is much more pronounced in gaster of males (Fig. 3c). Yellowish white, decumbent pubescence on the whole body, including scapes and legs. Longer erect, yellowish white hair especially on clypeus, but also on rest of head, alitrunk and gaster. Propodeum, seen in pro- file, broadly rounded and slightly higher than promesonotum. Dor- sal part of the propodeum convex, descending part straight. In the lower fifth, the declivity is slightly concave. Paratype $$ : TL 5.1-5.8, HL 1.26-1.38, HW 1.29-1.48, CI 98-106, SL 1.43-1.57, SI 85-100, PW 0.77-0.92, OD 0.31-0.39 or 0.24-0.31 H W (20 measured).
Holotype 2, Western Malaysia: Pahang, near Kuantan, 20.2.1984 (Naturhistorisches Museum, Basel).
Paratypes: 20 $$ with the same data as holotype (2 in British Museum (Natural History), London; 2 in Museum of Comparative Zoology at Harvard University; 2 in National Museum of Malaysia, Kuala Lumpur; 2 in Museo Civico di Storia Naturale, Geneva; 12 in collection of the author).
B. Gynetype(Q):TL 11.3, HL2.33, HW 2.24, CI96, SL2.16, SI 104, PW 1.76, OD 0.69.
Frontal carinae extend about to midlength of the head. Apart from a lateral projection beyond the scapal insertion, they are almost straight and only slightly divergent. Clypeus wider than that of worker (0.67 mm as against 0.50 mm). Its anterior margin shows a median semicircular excision, with indented edges (Fig. I b). Mandibles strong, distinctly rounded on outside and with 5 sub- equal teeth on inside. Frontal area not clearly delimited and hardly more shining than surroundings. Head and alitrunk black, tergites



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Fig. 1.
Scanning micrograph of Camponofus [exens: a, head of worker; b, head of female; c, alitrunk of worker in profile. of gaster dark brown; sternites lighter brown; front of head, distal antenna1 flagellum and legs yellow brown. Wings yellowish, veins yellow brown.
Head, alitrunk and gaster covered with fine, light yellow, decum- bent pubescence. Longer erect hairs distributed over the whole body. Eyes distinctly larger than those of workers and situated behind the midlength of the sides of the head. OD 0.69 mm, or about 0.30 HW. Petiole with a broad base, tapering toward the apex into a narrow ridge, seen from the side.



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Fig. 2.
Scanning electron micrograph of Componotits texens male: a, head; b petiole of alitrunk and gaster in profile. Paratype $9: TL 10.8-11.3, HL 2.29-2.46, HW 2.20-2.40, CI 95-98, SL 2.12-2.19, SI 101-107, PW 1.72-1.92, OD 0.69,ODj H^ 0.28-0.3 1 (4 measured),




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IYÌö~ uumpen - ~amponoiusjrom ~aiaysia 303 Gynetype, Western Malaysia: Gombak Valley, ca. 25 km north of Kuala Lumpur, 3 1.12.1982, U. Maschwitz leg. (Naturhistorisches Museum, Basel).
Paratypes: 4 $9 with the same data as gynetype (collection of the author).
C. Allotype ((5): TL 6.7, HL 1.0, HW 1.02, CI 102, SL 1.00, SI 102, PW 1.32, OD 0.46.
The trapezoidal head nearly as wide as long (CI 102), with promi- nent convex eyes, extending to the upper end of the head sides. Occipital margin strongly convex with protruding ocelli (Fig. 2a); clypeus narrow (width about 0.17 mm) with straight anterior mar- gin. Short frontal carinae sinuate, reaching back to midlength of head. Eyes very large; maximum diameter 0.46 mm, or about 0.45 HW. Scapes long, projecting beyond occipital margin of the head by more than half their length. Pedicel expanded at its distal end and thicker than following flagellar segments. Head and alitrunk dark brown like scapes and legs; petiole and gaster lighter brown; antenna1 flagellum yellow brown. Wings yel- lowish with lighter brown veins. Decumbent pubescence on head and alitrunk thin, but denser on gaster; in addition, body covered with longer, erect hairs. Cuticular sculpture of head, alitrunk and petiole reticulated, that of gaster imbricated (Fig. 3c). Propodeal profile rounded, with weakly convex dorsal and weakly concave descending part. Petiolar scale triangular in profile, with a broad base tapering to a ridge (Fig. 2b). Ridge with slight median excision. Paratype TL 6.5-6.7, HL 0.86-0.92, HW 0.93-0.99, CI 96-108, SL 1.00, SI 93-99, PW 1.16-1.24, OD 0.43-0.46 (4 mea- sured). Allotype, Western ~ala~sia: Gombak Valley, ca. 25 km north of Kuala Lumpur, 3 1.12.1982, U. Maschwitz leg. (Naturhisto- risches Musaeum, Basel).
Paratypes: 4 males with same data as allotype (collection of the author).
Camponotus gombaki sp. n.
Derivatio nominis: The name is derived from the Gombak Val- ley, in which the first and only colony of this species was found, and in honor of the 'Gombak Field Station of the University of Malaya,' where we worked during our research stay.



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19851 Dumpert - Camponotus from Malaysia 565 A. Holotype 9: TL 6.1, HL 1.46, HW 1.49, CI 102, SL 1 S6, SI 96, PW 0.93, OD 0.34.
Frontal carinae slightly sinuate, reaching back to midlength of the head (Fig. 4a). Anterior clypeal margin almost straight, but in the middle third weakly concave. Eyes situated behind the midlength of the sides of the head, with maximum diameter of 0.34 mm or about 0.23 HW.
Head and alitrunk reddish brown, gaster dark brown, almost black. Petiole reddish brown at base, but dark brown in upper half. Legs and antennae, like head and alitrunk, reddish brown to top. Surface of head, alitrunk and gaster opaque, with very dense punc- tures, which appear elongated on the gaster. Cuticular structure under SEM regular reticulated on head and irregular reticulated on alitrunk; cuticular sculpture of gaster is shown on Fig. 3a. Yellowish white, decumbent pubescence less dense than in C. texens workers; yellowish white, erect, longer hairs occur mainly on forehead and vertex, less dense on alitrunk and gaster. Propodeum in profile broadly rounded (Fig. 4c, d) and lower than promesonotal dorsum. Dorsal part of propodeum convex, descending part slightly flatter. In the lower sixth, the declivity is slightly concave. Paratype $9: TL 5.6-6.4, HL 1.40-1.62, HW 1.44-1.72, CI 97-105, SL 1.56-1.62, SI 86-104, PW 0.92-1.04, OD 0.34-0.40, or 0.22-0.26 HW (20 measured).
Holotype $, Western Malaysia: Gombak Valley, ca. 25 km north of Kuala Lumpur, 15.2.1984 (Naturhistorisches Museum, Basel). Paratypes: 20 QQ with same data as holotype (2 in British Museum (Natural History), London; 2 in Museum of Comparative Zoology at Harvard University; 2 in Malaysia, National Museum of Malay- sia, Kuala Lumpur; 12 in collection of the author). B. Gynetype (9): TL 11.4, HL 2.18, HW 2.24, CI 103, SL 2.38, SI 94.1, PW 1.92, OD 0.54.
Frontal carinae extend about to midlength of head. Apart from a lateral projection behind the scapal insertion, they are almost straight and hardly divergent. Width of clypeus subequal to that of C. texens. Anterior clypeal margin with semicircular excision (Fig. 4b). Mandibles strong, distinctly rounded on outside, and with 5 subequal teeth on inside. Frontal area clearly delimited and more shining than neighboring head parts.




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566 Psyche [VOI, 92,
Fig. 4.
Scanning electron micrograph of Camponotus gombuki; a, head of worker; b, head of female; c, alitrunk of worker in profile; d, petiole of worker, with parts of alitrunk and gastcr in profile. Head, alitrunk, petiole and legs reddish brown; gaster, antennae and mandibles dark brown. Wings yellowish, veins yellow brown. Body covered with short, yellowish white, decumbent pubescence; longer, erect, yellowish white hair is particularly dense on clypeus, but also on rest of body, including scapes and legs. Longer hair on alitrunk is less dense than in C. texens female. Eyes are situated behind the midlength of the sides of the head and markedly larger



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19851 Dumpert - Camponotus from Malaysia 567 than those of workers, but smaller than those of C. texens females; maximum diameter 0.54 mm, which is about 0.24 HW. Petiolar profile with broad base, which tapers to a transverse ridge and into a point, as seen from above.
Paratype @: TL 11.4, HL 2.21-2.23, HW 2.24-2.29, CI 103-104, SL 2.3-2.4, SI 94-98, PW 1.92-1.99, OD 0.58-0.62, OD/HW 0.24-0.25 (4 measured).
Gynetype, Western Malaysia: Gombak Valley, ca. 25 km north of Kuala Lumpur, 15.2.1984 (Naturhistorisches Museum, Basel). Paratypes: 4 QQ with same data as gynetype (collection of the author).
Camponotus overbecki Viehmeyer stat. n.
Camponotus dolichoderoides var. overbecki Viehmeyer 19 15: 162. Material examined: 1 syntype worker from the Santschi Collec- tion (Naturhistorisches Museum, Basel) leg. Overbeck: Singapore. TL5.8, HL 1.43, HW 1.56, CI 109, SL 1.61, SI97, PW0.88, OD 0.39.
Frontal carinae extend to about midlength of the sides of the head; as in C. texens, they are slightly divergent and-apart from a lateral projection behind the scapal insertion-straight. Anterior clypeal margin almost straight, but slightly concave in the middle third. Eyes are situated behind the midlength of the sides of the head; maximum diameter 0.39, which is about 0.39 HW. Head and alitrunk reddish brown, gaster blackish brown with lighter anterior segment boundaries. Petiole, legs, tarsi, and apical antenna1 flagellum yellow. Head, alitrunk and gaster shining, less punctured than in C. texens and in particular in C. gombaki. Whole body covered with fine, yellowish white, decumbent pubescence; longer erect denser than in C. texens and in particular in C. gom- baki. Propodeum in profile broadly rounded and slightly higher than promesonotal dorsum. Dorsal part of propodum weakly con- vex, descending part slightly flatter. In the lower fourth, the decliv- ity is slightly concave.
On the whole, the differences between C. gombaki and C. texens are not exceeding those between C. overbecki and C. dolichode- roides. As C. texens and C. gombaki certainly belong to different species because of the differences in their behavior (Maschwitz et



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568 Psyche [vo~. 92
al., 1985), there is no sufficient reason to continue designating C. overbecki as a variety of C. dolichoderoides. It is therefore made a species on its own. The female of C. overbecki is described by Vieh- meyer (1915) from an isolated specimen as follows: head narrower and more elongated than that of worker; eyes larger and more con- vex than those of workers and situated in front of the midlength of the sides of the head. Clypeus slightly emarginate anteriad. Dispersed, decumbent pubescence on the whole body. Female deal- ate and 6.5 mm long.
Body length of 6.5 mm is far smaller than that of any other known Karavaievia females. Head deviates markedly in shape and size, and eyes are situated in front of the midlength of the sides of the head and more convex than those of workers. This was not found in any other Karavaievia female. In contrast to all other Karavaievia females, clypeus slightly carinated and emarginated. These basic differences from all known Karavaievia females suggest that the female described by Viehmeyer is not a female of C. overbecki. Males of C. overbecki as described by Viehmeyer (1915): Head with strongly convex eyes slightly broader than long, occipital mar- gin strongly convex, anterior clypeal margin straight, mandibles with a large apical tooth. Propodeal profile with weakly convex dorsal and concave descending face; dorsal face with shallow, median, longitudinal furrow. Petiolar scale lower and slightly thicker than that of worker, tapering to a transverse ridge, which is slightly excised in the middle. Densely reticulated, front part of the body more opaque than gaster. Dirty yellow brown, gaster with indistinct narrow, dark bands in front of the posterior segment borders, femora and tibiae slightly darkened; wings yellowish with yellow brown veins. 6 mm in length.
Camponotus dolichoderoides Fore1
Camponotus dolichoderoides Forel, 19 1 1 : 5 1. (Types not found, presumed lost.) Description is translation of Fore1 (191 1): workers 6 mm in length; strong, curved, shining mandibles, with 5 teeth and widely separated punctures. Clypeus without carina, anterior margin straight, and not or only weakly protruding. Frontal carinae little sinuate and hardly divergent. Scapal insertion close to the posterior clypeal margin. Head trapezoidal, with strongly convex sides, and nearly as



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19851 Dumpert - Camponotus from Malaysia 569 broad as long with maximum diameter in the posterior third. Occipi- tal margin moderately concave; eyes situated close beyond the midlength of the sides of the head. Scapes project behind the occipi- tal margin by about one third of their length. Apical antennal flagel- lum slightly thickened. Promesonotal dorsum broadly rounded with deep promesonotal suture. Deep impression between promesono- turn and propodeum, with two raised stigmata on the ground. Dor- sal propodeum in profile slightly convex, forming almost a right angle with declivity which is almost straight. Petiolar scale in profile cone shaped. Gaster oval, tibiae cylindrical, not thickened and without spines. Cuticular surface weakly reticulated and slightly opaque, covered with punctures which are particularly dense and elongated on gaster. Decumbent pubescence very fine, yellowish, distributed over the whole body, including tibiae and scapes, espe- cially dense on gaster. Uniformly dirty yellow brown; tarsi, apical antennal flagellum and front head reddish to yellow red. Hayvep, Borneo.
Carnponotus exsectus Emery
Camponotus exsecrus Emery, 1901 : 53. Syntype Q$ and $9: Museo Civico di Stork Naturale, Geneva (examined).
Original description (Emery, 190 1): workers dirty yellow, shining, front head and gaster lighter, segment borders brownish, decumbent pubescence denser on head and alitrunk than on gaster. Body, including scapes and legs, covered with longer, whitish, erect hairs. Weak cuticular punctures on the whole body, those of the gaster elongated. Head rounded and convex. Clypeus weakly convex and not carinated; anterior clypeal margin with wide excision in the middle third, and indented edges. Mandibles strongly curved on outside and 5 teeth on inside. Frontal carinae almost straight, linea frontalis weak, but clearly visible. Scapes project beyond occipital margin by about one third of their length. Viewed in profile, pro- and mesonotum form a curved line which is connected to the pro- podeal outline by an obtuse angle. Propodeal profile broadly rounded. Scale biconvex, tapering into a sharp transverse ridge. Length: 4-314 to 5-1 /2 mm.
According to my own examination, workers are more shining than those of C. texens, but not as much as those of C. gombaki.



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570 Psyche [Vol. 92
Erect hairs more numerous than in C. texens. Propodeal dorsum clearly higher than dorsum of promesonotum, and in profile more inclined than in C. texens, C. gombaki, and C. overbecki. Dorsal part of propodeal profile convex, descending part nearly straight. Frontal area clearly delimited.
Original description of female (Emery, 1901): darker than workers; sides, scutellum, propodeal dorsum, legs, and major part of gaster brown. Head and mesonotum opaque; pubescence very dense, in particular on gaster. Head trapezoidal, mandibles similar to those of workers; clypeus narrow with deep semicircular excision in the middle of the anterior margin, limited by protruding acute angles. Antennae strong, petiole similar to that of workers. Punc- tures on gaster weak and not elongated. Length 11 mm. Mentawei, Sipora.
According to my own examination, C. exsectus females are larger than females of C. gombaki and C. texens. HW 2.5, HL 2.5, PW 2.3. Frontal carinae less divergent than those of C. texens and C. gom- baki females. Frontal area indistinctly delimited; mandibles reddish brown with black teeth; gaster shining.
Comparison of Karavaievia Species
The subgenus is characterized by morphological traits and possi- bly by behavioral characters, too. The common morphological traits include the subuniform size of workers, females and males, the shape of the head, the position of the eyes, the characteristic shape of frontal carinae, clypeus, mandibles, antennae and alitrunk. All known Karavaievia species originate from the Indo-Malayan area, in particular from Malaysia and Singapore. A possibly common behavior of all Karavaievia species is the weaving of silk nests on the undersides of leaves which, however, has only been investigated for C. texens and C. gombaki (Maschwitz et al., 1985). For C. over- becki, there is only the statement of Viehmeyer (1915) that Overbeck found workers and males in 'carton nests' on the underside of leaves. Nothing has become known so far, however, on the nest-building of C. dolichoderoides and C. exsectus.
My examination of two workers and a male of C. texens and of two workers of C. gombaki showed no openings of metapleural glands. This is in accordance with the observations in all weaver ants studied by Holldobler and Engel-Siege1 (1984).



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19851 Dumpert - Camponotus from Malaysia 571 The two species described by Fore1 (191 1) as C. horrens and C. moeschi, found on the Philippines, were assigned by Fore1 (19 1 1) to the same group as C. dolichoderoides, and also Chapman and Capco (1951) classified C. horrens in the subgenus Karavaievia. Emery (1925), on the other hand, classified C. moeschi in the subge- nus Myrmamblys and C. horrens in the subgenus Colobopsis; in the latter case, however, Emery was not quite sure, because neither soldiers nor females of this species had been found. C. horrens and C. moeschi definitely do not belong to the subgenus Karavaievia. This is supported in the case of C. horrens by the carinated and twice-excised anterior clypeal margin of the workers, the strongly divergent frontal carinae and the position of the eyes in the posterior fourth of the head. Features against C. moeschi as a member of the subgenus Karavaievia are the notched clypeus, lobed in the anterior part; the elongated head, convex in the occipital part; position of the eyes at the posterior third of the head, and the weakly impressed metanotal groove.
The differences between workers of the Karavaievia species are in coloration, sculpture and pubescence, slight differences in the shape of the frontal carinae, differences in size of head and eyes, and in shape and size of propodeum. A broader comparison of the Kara- vaievia sexuals, however, is not yet possible, because females of only 3 species and males of 2 species are known. Karavaievia workers are dirty yellow and brown, from reddish brown to blackish brown; in most cases, the front of the head, the apical flagellar segments and the tarsi are of a uniform color, which differs from the color of the rest of the body.
Key to the species of Karavaievia
Workers
1. Head and alitrunk shining and covered with erect hairs. ..... .2 Head and alitrunk opaque and only sparsely covered with erect hairs .................................................. 3 2. Uniformly dirty yellow, anterior parts of head and gaster lighter. Gaster with darker segment borders. Frontal carinae straight and diverging.. ................................... C. exsectus Head, alitrunk and legs reddish brown, gaster blackish brown with lighter brown segment borders. Apical antenna1 flagellum



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572 Psyche [vo~. 92
and front of head yellow. Frontal carinae weakly sinuate ...... ............................................ C.overbecki 3. Propodeal profile arched, dorsal and descending parts almost straight, forming an angle of about 90'. Uniformly dirty yellow- ish brown; tarsi, apical antennal flagellum and front of head ..............................
reddish.. C. dolichoderoides
Propodeal profile rounded .............................. .4 4. Propodeurn higher than promesonotum. HW 1.35- 1.5 1, HL 1.3 1 - 1.47, OD 0.24-0.28 H W. Head, alitrunk, scapes and legs dark brown; gaster, petiole, apical antennal flagellum and front ............................
of head dirty yellow.. C. texens
Propodeum lower than promesonoturn. HW 1.44- 1.72, HL 1.40-1.62, OD 0.22-0.26 HW. Head and alitrunk reddish brown at base, and darker brown in upper half. Head and alitrunk .................
opaque and densely punctured.
C. gombaki
Females
1. Petiolar scale blunt in dorsal view. ............... C. exsectus .......................
Petiolar scale acute in dorsal view.
.2
2. Head, alitrunk, petiole and legs reddish brown, gaster, antennae and mandibles dark brown. Frontal area clearly delimited and more shining than the surroundings. Gaster opaque. OD ....................
0.54-0.62, or 0.24-0.25 HW C. gombaki
Head and alitrunk black, tergites of gaster dark brown, sternites brown. Frontal area clearly delimited. Gaster shining. OD 0.69 ...............................
or 0.28-0.31 HW.. C. texens
Males
1. Color on the whole dirty yellowish brown with indistinct narrow bands in front of the posterior segment borders .............. .......................................... C. overbecki Head and alitrunk dark brown like scapes and legs; petiole and gaster lighter. Antenna1 flagellum yellowish brown ........... .......................................... C-gombaki



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Two new silk nest weaving species, Camponotus texens and Camponotus gombaki, are described as members of the subgenus Karavaievia Emery 1925. The subgenus now consists of five species , which are probably all weaver ants. Additionally, a key to the spe- cies of Karavaievia is presented.
BOLTON, B.
1977. The ant tribe Tetramoriini (Hymenoptera: Formicidae). The genus Tetramorium Mayr in the Oriental and Indo-Australian regions, and i n Australia. Bull. Brit. Mus. (Nat. Hist.), Ent., 36(2): 67-151. CHAPMAN, J. W. & S. R. CAPCO
1951.
Check list of the ants (Hymenoptera: Formicidae) of Asia. Manila Bureau of Printing.
EMERY, C.
1901.
Formiche racolte da elio Modigliana in Sumatra, Engano e Mentawei - Ann. Mus. Stor. Nat. Genova, 40: 709-743. 1925.
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