C. H. T. Townsend.
On the Tribe Dejeaniini of the Muscoid Family Hystriciidae, with Five New Genera.
Psyche 20(3):102-106, 1913.

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108 Psyche [June
ON THE TRIBE DEJEANIINI OF THE MUSCOID FAMILY HYSTRICIIDB, WITH FIVE NEW GENERA.
BY CHARLES H. T. TOWNSEND,
Director of Entomological Stations, Lima, Peru. The Dejeaniine flies are among the most striking and interesting of the muscoid group.
They are characterized not only by a
general strong macrochsetal development showing in more or less densely placed spines on the abdomen and scutellum, which they share with many other forms of the Hystriciidce, but also by the unique development of the palpi which are greatly elongated. The tribe as so far known to the writer divides naturally into the four subtribes ~ejeaniiia, Paradejeaniina, Lasiopalpina and Dejeaniopina, the last three being monotypic. Only the last two have pilose eyes. The Dejeaniina divide into three group-units, the Dejeaniioe with discal spines on intermediate abdominal seg- ments and normal palpi, the Eulasiopalpioe without discal spines and with ciliate palpi, and the Eudejeaniice without discal spines and with normal palpi, Dejeania was founded by Robineau Des- voidy in 1830, who included therein at the time the two species bradiensis RD, and capensis RD., but without designating either as the type. In 1910 Coquillett designated the latter, which equals Stomoxys bombylans F., as type of the genus. Lasiopalpus was founded by Macquart in 1847, with the single species flavitarsis Mcq. which is thus the type. Eudejeania was founded by the writer in 1912, with designation of subalpina T. as type. All the Dejeaniine flies known up to twenty years ago, other than Lasiopalpus, were grouped under the single genus Dejeania. Brauer and von Bergenstamm were the first to cut any of the forms loose from this combination, characterizing the genus Parade- jeania in 1893 for Dejeania rutilioides Jaennicke and Jurinia myrrhea Say, BB. As the latter is a momen nudum, the former stands as the type of the genus. In 1898 Brauer added to the genus, Jurinia hystrix Riley (nee F., nec Will.) MS from North America, Jurinia amethystina Macquart from Brazil (Minas- Gerags), Jurinia bicolor Macquart (nec Wd.) from Colombia and Cayenne. (two species, probably MS in coll.), Jurinia nigrwdlyp-



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19131 Townsend-On the Tribe Dejeaniini of the Muscoid Family 103 trata Macquart from Brazil (Minas-Geraes), and Hystricia erythrina Bigot from Brazil (Bahia). The remainder of the forms continued under the old genus until 1912, when the writer established the genus Eudejeania for certain strikingly large and distinct forms which range high in the Andean montanya. The African Dejeania capensis ED. (equals bombylans l?., Wd.) is not congeneric with any of the American forms, hence its desig- nation as type of the genus leaves the latter without nearer generic reference than Eudejeania, which is strikingly contrasted with the older and better known forms. The figure 21, on page 44 of the third edition of Dr. Williston's Manual, labeled Paradejeania sp., is evidently not that genus. It has almost exactly the form and aspect of Eudejeania female; if discal spines are wanting on the intermediate abdominal segments (they appear to show on the left side of second segment), and the probosjcis and palpi are both very elongate, it is probably Eudejeania. The figure 60, on page 366 of same work, also labeled Paradejeania sp., is neither Para- dejeania nor Eudejeania, but has the palpal characters of Jurinia. Tachina armta Wiedemann was taken by Brauer and von Bergenstamm in 1889 as typical of the ~merkan forms, and is here made the type of the new genus Adejeania. Tachina corpu- lenta Wiedemann and Dejeania vexatrix Osten Sacken are strikingly contrasted with Adejeania and the other American forms, the second showing an approach to Servillia in its long dense pubes- cence and the first exhibiting bristles in place of this pubescence. Both are made the types of new genera. Eulasiopalpus is erected for forms with ciliate palpi, long proboscis and bare eyes. The following synoptic table will show the distinctions between all of these forms with greatly elongated palpi, including the new genus Dejeaniops Townsend whose characterization is given in full in a forthcoming paper on the Hystriciidoe of the Andean montanya. Table of Tribe Dejeaniini.
1. Eyes thickly pilose, third antennal joint straight on front border and widened- ............................................... subtruncate apically. .2
Eyes practically bare, third antennal joint more or less convex on front border ........................................ and always rounded apically. .3 2. Palpi long and narrow, of even width except on the more narrowed base, not ciliate; proboscis elongate and slender; parafacials hairy; abdomen subovate and gently emarginate anally, no discal spines on intermediate segments; front



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104 Psyche [June
tarsi of female not conspicuously widened (Dejeaniopina) Dejeaniops gen. nov.
Palpi much widened on apical half, gradually narrowing basally, densely ciliate with long hair; proboscis short and stout; parafacials bare; front tarsi of female strongly widened (Lasiopalpina) .................... Lasiopalpus Mcq. Palpi narrow, gradually narrowing basally, ciliate with long hair; proboscis slender and conspicuously longer than head-height; parafacials hairy; abdo- ... men emarginate anally (Dejeaniina: Eulasiopalpi~e) Eulasiopalpus gen. nov.
Palpi not ciliate. ..................................................... .4 Palpi gradually widened from base distally, proboscis short; abdomen sub- quadrate and conspicuously emarginate anally; front tarsi of female strongly widened (Paradejeaniina) ...................... Paradejeania B. & von B. Palpi of even width on distal two-thirds or more; proboscis elongate and slender (Dejeaniina: Dejeaniise and Eudejeaniise) ............................. .5 Discal spines on dorsum of intermediate abdominal segments (Dejeaniise) Dejeania R.D. (Africa)
No discal spines (Eudejeaniiae) ........................................ .6 Abdomen subquadrate and deeply emarginate or buttocks-like behind in both sexes; front tarsi of female widened. ....................... Eudejeania T. Abdomen subovate, not broadly buttocks-like behind, at most the hind border of anal segment half as long as greatest width of abdomen and gently emargi- ..................... nate in female; front tarsi of female not widened. .7
Abdomen clothed with strong bristles in addition to the spines and differentiated therefrom, front tarsi of male with bristly hairs on inside Echinotachina gen. nov.
.................. Abdomen with only pubescence in addition to the spines. .8 Pubescence short and normal. ....................... Adejeania gen. nov. Pubescence long, dense, even and pronounced, showing conspicuously among the spines, thorax also with a dense pubescence. ... Trichodejeania gen. nov. Below is a statement of the genera and their types. Whether
all these forms belong in one natural tribe remains to be decided by a careful intensive study of the group and its allies. Although it would seem probable that these elongate palpi are not all of common origin, yet none of the forms exhibits any striking depart- ure in its external anatomy from the general characteristics of the group. A comparative study of the first-stage maggots, which the writer has in hand, will conclusively demonstrate the actual affinities.
DEJEANIA R. D. (1830)-Type, Stomoxys bombylans F. (equals Tachina bombylans Wd., and Dejeania capensis R.D.) . LASIOPALPUS Mcq. (1 847)-Type, L. flavitarsis Mcq. PABADEJEANIA B.B. (1893)-Type, Dejeania rutilioides Jaen. EUDEJEANIA T. (1912)-Type, E. subalpina T.



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19131 Townsend-On the Tribe Dejeaniini of- the Muscoid Family DEJEANIOPS T. gen. nov.-Type, D. ollachea T. n. sp. EULASIOPALPUS gen. nov.-Type, Lasiopalpus albipes T. n. sp. ADEJEANIA gen. nov.-Type, Tachina armta Wd. (equals De- jeania armata BB.).
TRICHODEJEANIA gen. nov.-Type, Dejeania vexatrix O.S. ECHINOTACHINA gen. nov.-Type, Tachina corpulenta Wd. The following descriptions are abridged from the originals which will appear in the forthcoming paper on the Hystriciidce of the Andean montanya.
Dejeaniops ollachea sp. nov.
Length, 11 mm. One female, OHachea, San Gaban canyon, southeastern Peru, about 9,500 ft., Feb. 2 (Townsend). Head yellowish, faintly golden. Palpi rust- yellow, antennse yellowish-brown to reddish, arista black. Parafrontals and thorax
brownish-olive, scutellum pale. Abdomen soft black, with clear light yellow on sides of anterior two-thirds. Legs rust-yellow, femora largely blackish. Wings and tegulse smoky, tegulse thickly microscopically black-pubescent. Palpi and proboscis both very long and slender, parafacials and eyes pilose. Lasiopalpus albipes sp. nov.
Length, 14 mm.
One male, Huascaray ridge, northern Peru, about 7,000 ft., Sept. 22 (Townsend). Head smoky-silvery; epistoma, front, antennae and palpi black or blackish. Thorax, scutellum, abdomen and femora black, tibias pale yellowish, tarsi yellowish-white. Wings infuscated, tegulse smoky-black. The third antennal joint is rounded apically, eyes bare, parafacials hairy, and proboscis about one and one-fourth times head-height. The following are descriptions of two new species of Eudejeania recently secured by the writer in the high Andean region of central Peru :-
Eudejeania punensis N. sp.
Length of body to ends of spines, 14 to 16 mm.; of wing, 11.5 to 13.5 mm.; of palpi, 3.5 to 4 mm.; width of abdomen, 7 to 8 mm. Fifty-two females and one male, Pachacayo valley, about 12,000 feet, in puna region about half way between Oroya and Jauja, Peru, March 27 and 28, 1913, on flower's of Viguiera-like composite. The immense preponderance of the females is worthy of remark. Differs from Eudejeania nigra T. by its much smaller size and by the following colorational characters:-Legs with their spines, palpi and first two antennal joints rust-yellow, the second antennal joint more or less slightly smoky on base. Wings only lightly smoky, the veins rust-yellow giving the wing bases a distinct yellow tinge. Parafrontals dusky olive. Occipital pile pale brassy. Thorax and abdo- men dark brown to black, showing smoky pollinose on anal segment in oblique view and somewhat less so on third segment.



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106 . Psyche [June
Type, TD4135 (female fly, reproductive system, first-stage maggot). Has strap-like uterus and deposits blackish maggots, probably on foliage. Eudejeania alpina n. sp.
Length of body to ends of spines, 20 mm.; of wing, 16.5 mm.; of palpi, 5 mm.; width of abdomen, 11 mm. Two females, Pachacayo valley, about 12,000 feet, with preceding species on same flowers, March 28, 1913. Differs from Eudejeunia punensis T. by its much larger size and by the following colorational characters :-Head deeply fuscous, practically black, only narrow orbits and wider border in front of eyes brassy. Antennae wholly black. Frontalia
silvery to tawny pollinose in oblique view. Palpi yellowish or tawny, but deeply tinged with smoky. Occipital pile golden to brassy. Wings deeply smoky through- out including bases; veins brown to blackish, not showing yellowish to the unaided eye.
Legs slightly deeper rust-yellow, the femora more reddish. Type, TD4139 (female fly, reproductive system). It should be noted here that the writer, in all of his collecting, has never before found such an abundance of Dejeaniine flies in the height of activity as that encountered at the above-mentioned locality in Pachacayo valley, forty-five of the above specimens having been taken there in an hour or so of fitful sun and rain during the forenoon of March 28. Hundreds of E. punensis could doubtless have been collected in the locality had sun and time permitted. An approach to such abundance, but in a state of dulled activity, was encountered in October, 1895, in the Sacra- mento Mountains of southern New Mexico (Ann. & Mag. N. IL, 6th ser., vol. 19, No. 110, pp. 144-145).



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